Loboclina Grishin, 2019

Grishin, Nick V., 2019, Expanded phenotypic diagnoses for 24 recently named new taxa of Hesperiidae (Lepidoptera), The Taxonomic Report of the International Lepidoptera Survey 8 (1), pp. 1-16 : 3-4

Subtribe Loboclina Grishin, 2019

Type genus: Lobocla Moore, 1884 .

ZooBank registration: C606FC35-323D-4E55-AF5A-A86C6366BAFA

Diagnosis: The subtribe was defined as a clade from about 30 Mya that contains the genus Lobocla , and putatively synapomorphic DNA characters for this clade were given ( Li et al., 2019). Phenotypically, species in this subtribe would key to B. 4 in Evans (1949) or C.5, C.10a, C.15.2 or C. 18 in Evans (1953). Genera in this tribe are unified by the long forewing cell, at least 2/3 of costa, lobed or tailed hindwing, short, non-divergent palpi with the 2nd segment touching the face and the 3rd not protruding beyond the 2nd (except Zestusa ). Most genera could be diagnosed by single character and possible synapomorphy: broadly arcuate antennal clubs ( Fig. 2 ). Additionally, hyaline spot in forewing cell R 2 -R 3 present even in species with hooked antennae. In species with hooked antennae that lack the spot, genitalic valvae asymmetrical and very broad. Typically (except Aguna ), valvae rather broad, somewhat rectangular, with harpe narrow, long upturned, hook-like, leaving a deep space between harpe and ampulla, uncus divided.

Genera included: Aguna R. Williams, 1927 , Zeutus Grishin, 2019 , Lobocla Moore, 1884 , Lobotractus Grishin, 2019 , Codatractus Lindsey, 1921 , Zestusa Lindsey, 1925 , Ridens Evans, 1952 , and Venada Evans, 1952 .

Parent Taxon: Tribe Eudamini Mabille, 1877 .

Type genus: Cephise Evans, 1952 .

ZooBank registration: 22B59811-F174-4FDF-A9D2-799897F4D44E

Diagnosis: The subtribe was defined as a clade from about 30 Mya that contains the genus Cephise , and putatively synapomorphic DNA characters for this clade were given ( Li et al., 2019). Phenotypically, species in this subtribe would be diagnosed by "genitalia and palpi as described by Burns (1996: 182-183) for Cephise " ( Li et al., 2019). Note that this statement refers to the characters of genitalia and palpi (they are "as described... for Cephise "), not to the genus Cephise . Thus, the referenced publication could have been describing the characters of any other genus. Not the genus is relevant here, but the characters themselves, some of which are the following: uniquely short, truncate uncus with a couple of teeth at each distal corner, harpe extending distad, separated from the valva by a U-shaped notch; the first segment of palpi below the eyes with a shelf-like projection of scales that looks like a triangle below the eyes ( Fig. 3 ), more developed in males than in females.

Genera included: Cephise Evans, 1952 .

Parent Taxon: Tribe Eudamini Mabille, 1877 .

Figures

Fig. 2. Loboclina. Above: arcuate antennal club of Lobocla proximus (Leech, 1891), China: Yunnan Province. Below: hooked antennal club of Aguna coeloides Austin & O. Mielke, 1998, Brazil: Para.

Fig. 3. Cephisina. Head of Cephise cephise (Herrich-Schäffer, 1869) ♂, ventral view showing lateral projections of scales from palpi near eyes (by antennae).