Elymus transhyrcanus (Nevski) Tzvelev

Alieva, Kumush B., Peng, Yilong, Usupbaev, Adilet, Tojibaev, Komiljon Sh., Yusupov, Ziyoviddin, Ergashov, Ibrokhimjon, Azimova, Dilnoza & Jiang, Zhilin, 2025, Synopsis of the genus Elymus (Poaceae) in Uzbekistan (Middle Asia) with a description of Elymus uzbekistanicus a new species from Turkestan Mts, PhytoKeys 257, pp. 9-50 : 9-50

9. Elymus transhyrcanus (Nevski) Tzvelev in Novosti Sist. Vyssh. Rast. 9: 61 (1972)

≡ Agropyron transhyrcanum (Nevski) Bondarenko in Opred. Rast. Sred. Azii 1: 173 (1968). ≡ Roegneria transhyrcana Nevski in Trudy Sredne-Aziatsk. Gosud. Univ., Ser. 8 b, Bot. 17: 70 (1934).

= Elytrigia vvedenskyi Drobow in Fl. Uzbekistan. 1: 539 (1941). Type. Uzbekistan. Chulbair Mountains. Valley of the Obi-Dara River near the village of Sina. 31 V 1929, Vvedensky, 184.

= Roegneria leptoura Nevski in V. L. Komarov (ed.), Fl. URSS 2: 623 (1934). Type. Turkmenistan. Mountains of Chapan-Dag (Kopet-Dag). Collector unknown (holotype LE). ≡ Semeiostachys leptoura (Nevski) Drobow in Fl. Uzbekistan. 1: 285 (1941).

Type.

Turkmenistan • Ashgabat District, rocky areas at an elevation of 1000 m, Mountain Chapandag , 25 VIII 1931, No. 725, A. Borisova (isotype LE) .

Description.

Stems 80–95 cm, glabrous, smooth. Leaves 1.4–4 mm wide, grayish green, flat, margin rolled, adaxially rigid, abaxially slightly rough or glabrous. Sheath glabrous, smooth. Ligule to 1 mm. Spikes upright, rarely slightly drooping. Spikelets solitary along axis of spikes. Internodes of spike short hairy. Glumes almost equal, narrowly lanceolate, acuminate, with awn to 3 mm. Lemma lanceolate or narrowly lanceolate, slightly rough or glabrous, apex acute, with awn to 2 mm. Palea narrowly lanceolate, margins short ciliate. Anthers 3.5–4.5 mm (Fig. 11 ). 2 n = 56 ( Tzvelev 1976).

Phenology.

Flowering and fruiting: June-August.

Habitat.

On rocky slopes, cliffs, and pebbles in middle and upper mountain belts, 1700–2400 m.

General distribution.

Eastern and southern Transcaucasia, Iran, Turkey, Middle Asia (Tian Shan, Pamir-Alay: Hissar, Darvaz Ranges): Kazakhstan, Tajikistan, Turkmenistan, Uzbekistan.

Distribution in Uzbekistan.

Jizzakh, Kashkadarya, Surkhandarya, and Tashkent regions (Fig. 3 H ). I- 1 Western Tian Shan district. I- 1 - b Western Chatkal region. CHATKAL RANGE (Surroundings of the Chimgan Botanical Station. Big Chimgan, 07.1929, Gomolitsky s. n. [TASH 047581!]; Kashka-Su River Gorge, 26. 07. 1936, Korotkova, Titov 1702 [TASH 047578!]; Ridge between the upper Bashkizylsay and Chouli. Northern slopes, 29. 07. 1936, Korotkova, Titov 1847 [TASH 047579!]; Basin of the Chatkal River. Valley of the Akbulak River. South-Western slope of the first watershed ridge of the upper Akbulak River. Shrub and tall grass vegetation, 03. 09. 1938, Pyatayeva, Momotov 1564 [TASH 047570!]; Reserve, Chatkal Mountains, 70 km E of Tashkent 314199 [USDA _ NPGS 3436603], 314202 [USDA _ NPGS 3436605]; New section of Reserve, Chatkal Mountains, 70 km E of Tashkent 314206 [USDA _ NPGS 3436628], 314208 [USDA _ NPGS 3436624]); I- 1 - d Kurama region. KURAMA RANGE (Northern slopes. Near the Shaugaz Pass. Northern slope. Soil: fine-grained, brown, mountain-forest, 10. 07. 1939, Kudryashev 553 [TASH 046184!, TASH 046185!]). I- 5 Kuhistan district. I- 5 - a North Turkestan region. TURKESTAN RANGE (5–6 km south of the village of Tenga-Tapty, 19. 08. 1937, Demurina 1167 [TASH 047582!]; Northern slopes of the Turkestan Range, headwaters of the Sanzar River. Guralash-Say Nature Reserve. On the northeastern slope of Angyrly-Say, 26. 08. 1937, Korotkova, Vasil’kovskaya 1143 [TASH 047576!]). I- 6 Western Hissar district. I- 6 - a Kashkadarya region. HISSAR RANGE (Western Pamiro-Alay. Upper reaches of the Yakabagh-Darya River. Surroundings of the village of Tash-Kurgan. Kapyr-Say locality, 04. 07. 1936, Bochantsev, Butkov 625 [TASH 047574!, TASH 047575!]; Northern slopes of the Gissar Range. Basin of the Kashka-Darya River. Upper part of the Tanhas River basin. Headwaters of the Tanhas River, 01. 08. 1937, Kudryashev 1306 [TASH 047572!, TASH 047573!, TASH 047577!]; 02. 08. 1937, Kudryashev 1339 [TASH 055428!, TASH 055429!, TASH 055430!]); I- 6 - c Baysun region. HISSAR RANGE (Chulbair Mountains. Valley of the Obi-Dara River above the village of Sina, 31. 05. 1929, Vvedensky 184 [TASH 0000179!, TASH 0000180!]). I- 7 Hissar-Darvaz district. I- 7 - a Sangardak-Tupalang region. HISSAR RANGE (Pamiro-Alay. Gissar Range, Delli-Say, above the mouth of the Khucha River, elevation 2300 m. Description No. 87, 16. 08. 1941, Gromakov 729 [TASH 047583!]; Basin of the Tupolang River. Valley of the Khovat River. Stream on the right bank of the Khovat River, at the confluence of the Artushgar stream, 06. 09. 1947, Pyatayeva 483 [TASH 047583!]; Headwaters of the Shatrut River, upstream along the Chilik-Su River to the glaciers, and the surroundings of Chirmak-Zor, 28. 06. 1948, Pyatayeva 817 [TASH 047584!].

Figures

Figure 3. Distribution maps of Elymus species in Uzbekistan A E. caninus B E. lachnophyllus C E. fedtschenkoi D E. longearistatus E E. macrochaetus F E. nevskii G E. praeruptus H E. transhyrcanus I E. tschimganicus J E. tianschanigenus.

Figure 11. Elymus transhyrcanus a spike b spikelet c glumes d lemmas and paleas e stem f node g ligule (https://doi.org/10.13140/RG.2.2.14976.85763).