Clistopyga nubiphila Palacio & Bordera, 2025

Bordera, Santiago, Palacio, Edgard & Herrera-Flórez, Andrés Fabián, 2025, The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part VI: the C. calixtoi species group, with the description of twenty-four new species, Zootaxa 5662 (1), pp. 1-115 : 81-83

27. Clistopyga nubiphila Palacio & Bordera , sp. nov.

( Figs 41 , 51A , Appendix I)

Diagnosis. Clistopyga nubiphila may be distinguished from all other species of the C. calixtoi species group by the following combination of characters: genal orbits widely interrupted with dark brown or black ( Figs 41C–D ); mesoscutum reddish orange ( Fig. 41D ); pronotum bicoloured black and white cream ( Figs 41B, 41D ); mesopleuron entirely orange except the white cream subalar prominence ( Fig. 41D ); propodeum with lateral white cream or yellow marks ( Fig. 41D ); metapleuron bicoloured black and white cream ( Fig. 41D ); metasoma mostly orange to reddish brown without posterior defined white cream wide marks on tergites ( Figs 41E, 41J–K ); epomia absent ( Fig. 41B ); submetapleural carina absent ( Fig. 41D ); ovipositor relatively slender evenly and slightly up-curved at distal 0.35 ( Figs 41E–F ), about 2.3× as long as hind tibia ( Fig. 41E ); ovipositor sheath about 1.9× as long as hind tibia; ventral margin of ovipositor sheath basally with very small denticles.

Description. Female: Body length 11.5 mm. Fore wing length 7.9 mm.

Head ( Figs 41A, 41C–D ). In dorsal view, strongly narrowed behind eyes. Gena smooth and shiny with uniformly sparse setiferous punctures, in dorsal view almost straight, about 0.27× as long as eye, in frontal view more or less straight and moderately constricted below eyes. Frons and vertex smooth and shiny, with very isolated setiferous punctures. Posterior ocellus separated from eye about 1.04× its maximum diameter. Distance between hind ocelli about 0.75× maximum diameter of posterior ocellus. Hypostomal carina not lamelliform behind mandible. Face with fine and dense setiferous punctures, distance between punctures more than twice the diameter of punctures. Clypeal suture strongly curved. Clypeus about 1.53× as broad as medially long, distinctly convex in dorsal half, flat in ventral half, with ventral margin slightly concave. Malar space about 0.68× as long as basal mandibular width, with a slightly granulate subocular sulcus. Antenna with 32 flagellomeres, first flagellomere about 6.35× as long as wide.

Mesosoma ( Figs 41B–D, 41G–I ). Pronotum smooth and shiny with fine and scattered setiferous punctures in dorsal posterior part. Epomia absent. Mesoscutum entirely smooth and shiny, dorsally with fine setiferous punctures; anterior half of median lobe not conspicuously prominent, confluent with posterior half. Notaulus moderately deep, reaching about 0.5 the length of mesoscutum. Mesopleuron smooth and shiny, with evenly sparse setiferous punctures on anterior and ventral part. Epicnemial carina strong, its dorsal end slightly curved towards the back, ending at level of centre of pronotum. Metapleuron shiny, glabrous, except for some very sparse setiferous punctures at posterior part, about 2.87× as long as high. Submetapleural carina absent. Propodeum smooth and shiny, with evenly sparse fine setiferous punctures on anterior half and laterally, in dorsal view about 1.52× as long as medially wide. Propodeal spiracle about 0.16 the length to anterior end of pleural carina. Hind leg with femur about 4.0× as long as high, 1.10× as long as tibia. Fore wing with vein cu-a opposite to Rs & M; vein 2 rs-m about 0.58× as long as abscissa of M between 2 rs-m and 2 m-cu; abscissa of Cu 1 between 1 m-cu and Cu 1 a about 1.23× as long as Cu 1 b. Hind wing with vein cu-a about 0.33× as long as first abscissa of Cu 1 between M and cu-a, vein cu-a slightly reclivous, first abscissa of Cu 1 vertical and straight; distal abscissa of Cu 1 weakly pigmented.

Metasoma ( Figs 41E–F, 41J–K ). Tergite I about 1.9× as long as posteriorly broad, shiny, with sparse setiferous punctures laterally and posteriorly, spiracle near its anterior 0.4; lateromedian longitudinal carinae present at anterior 0.2 of tergite, very weak; lateral longitudinal carinae present at posterior 0.2 of tergite. Sternite I extending posteriorly about 0.5 the length of tergite. Tergite II 1.45× as long as posteriorly broad, with relatively dense and shallow setiferous punctures, distance between punctures about its diameter; remaining tergites same microsculpture. Ovipositor relatively slender, evenly and slightly up-curved at distal 0.35, about 2.3× as long as hind tibia. Ovipositor sheath about 1.9× as long as hind tibia, ventral margin basally with very small denticles, length of setae on average about 2.4× the sheath basal width.

Colouration ( Fig. 41 ). Head white cream with apex of mandibles, subocular sulcus, clypeal suture, a narrow longitudinal stripe at middle of face, frons and vertex medially, genal orbit medially and dorsally, posterior part of gena, and occipital region, black; antenna blackish brown, scapus, pedicel, annellus and first and second flagellomeres, laterally white cream. Mesosoma mostly reddish orange; dorsal part of propleuron, mid lateral part of pronotum, periphery of mesoscutum, axillae, ventral part of metapleuron, metasternum, propodeum extensively and groove separating propodeum from metapleuron, black; ventral part of propleuron, dorsal and anterolateral parts of pronotum, tegula, subalar prominence, mesepimeron, lateral parts of propodeum and dorsal part of metapleuron white cream. Metasoma with tergite I black with lateral margins marked with white cream; tergites II to VIII more or less entirely orange to reddish brown; tergites II–III black marked posterolaterally; ovipositor brown; ovipositor sheaths black.Anterior two pair of legs predominantly white, trochanters proximally black, femora and tibia dorsally striped with black and tarsi infuscate; hind leg white with coxa laterally, trochanter proximally, femur laterally and proximally, and tibia laterally, proximally, and distally black; hind tarsal segments predominantly blackish. Wings weakly infumate, pterostigma blackish brown.

Male: Unknown.

Etymology. The specific epithet nubiphila derives from the Latin “ nubes ” (cloud) and “ philus ” (loving) in reference to the Andean cloud forest where the type material was collected.

Type material (2♀♀). Holotype. Colombia : 1♀, Valle ,PNN Farallones de Cali , LosAndes,vda Quebradahonda, 3 º 34'N, 76 º 40'W, 1730 m, 1–8-IV-1998, malaise trap, N. Beltrán ( IAvH) . Paratype. 1 ♀, same locality, 25-XI-1998, W. Alfonso ( IAvH) .

Distribution. Colombia ( Fig. 51A , Appendix I).

Figures

FIGURE 41. Clistopyga nubiphila sp. nov., female (holotype): (A) head (frontal view); (B) epomia (lateral view); (C) head and mesosoma (dorsal view); (D) head and mesosoma (lateral view); (E) metasoma (lateral view); (F) ovipositor (lateral view); (G) fore wing; (H) fore wing; (I) hind wing; (J) metasoma (dorsal view); (K) tergite I and II (dorsal view). Scale bar: 2 mm (E); 1 mm (D, J, F); 0.5 mm (A, C, H, K); 0.2 mm (B, G, I).

FIGURE 51. Geographic distribution of C. calixtoi species group: (A) C. nasa sp. nov., C. nubiphila sp. nov.; (B) C. carvajali Gauld, Ugalde & Hanson, C. meridae sp, nov., C. stanfordi Gauld.