Clistopyga stanfordi Gauld, 1991

Bordera, Santiago, Palacio, Edgard & Herrera-Flórez, Andrés Fabián, 2025, The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part VI: the C. calixtoi species group, with the description of twenty-four new species, Zootaxa 5662 (1), pp. 1-115 : 91-94

32. Clistopyga stanfordi Gauld, 1991

( Figs 46 , 51B , Appendix I)

Clistopyga stanfordi Gauld, 1991: 299 . Holotype: ♀, Costa Rica (NHMUK).

Diagnosis. Clistopyga stanfordi may be distinguished from all other species of the C. calixtoi species group by the following combination of characters: genal orbits widely interrupted with brown or black ( Figs 46A, 46E ); mesopleuron ventrally white cream ( Figs 46A, 46E ); metasoma with posterolateral well defined white cream marks on tergites, clearly separated from posterior rim ( Figs 46A, 46I ); head in dorsal view, strongly narrowed posteriorly to eyes ( Fig. 46D ); gena in dorsal view straight, 0.20–0.25× as long as eye ( Fig. 46D ); epomia as a small tubercle or short carina at the bottom of the submarginal depression of pronotum ( Fig. 46C , arrow); metapleuron 2.15–2.2× as long as high ( Fig. 46F ); submetapleural carina extended on anterior 0.4 of metapleuron ( Fig. 46F ); hind leg with femur 4.3–4.5× as long as high; ovipositor very slender, slightly and evenly up-curved at 0.4 distal, 1.8–1.85× as long as hind tibia ( Figs 46A, 46J ); ovipositor sheath 1.45–1.5× as long as hind tibia.

Description. Female: Body length 6.4–9.7 mm. Fore wing length 4.6–6.2 mm.

Head ( Figs 46A–B, 46D–E ). In dorsal view, strongly narrowed posteriorly to eyes. Gena smooth and shiny with very sparse setiferous punctures, in dorsal view straight, 0.20–0.25× as long as eye, in frontal view strongly constricted ventrally to eyes, with almost straight profile. Frons and vertex smooth and shiny, with very isolated setiferous punctures. Posterior ocellus separated from eye 0.70–0.90 its maximum diameter. Distance between hind ocelli 0.7–0.9× maximum diameter of posterior ocellus. Hypostomal carina not lamelliform posteriorly to mandible. Face with shallow and moderately dense punctures, distance between them more than twice the diameter of punctures. Clypeal suture moderately curved Clypeus 1.65–1.75× as broad as medially long, moderately convex with isolated fine punctures dorsally, flat and smooth ventrally, ventral margin almost straight. Malar space 0.7– 0.85× as long as basal mandibular width, with shallow and slightly granulate subocular sulcus. Antenna with 26–28 flagellomeres, first flagellomere 6.0–6.7× as long as medially wide.

Mesosoma ( Figs 46A, 46C, 46D–H ). Pronotum smooth and shiny with few setiferous punctures on posterior dorsal corner. Epomia as a very short carina or tubercle at the bottom of the submarginal depression of pronotum. Mesoscutum shiny, with moderately dense setiferous punctures, denser on anterior part of median lobe; anterior half of median lobe not conspicuously prominent, confluent with posterior half. Notaulus weak, reaching about 0.5 the length of mesoscutum. Mesopleuron shiny, with evenly sparse setiferous punctures on anterior and ventral part. Epicnemial carina moderately strong, its dorsal end straight, ending at level of centre of pronotum. Metapleuron, smooth and shiny, glabrous, except for some sparse setiferous punctures at posterior part, 2.15–2.2× as long as high. Submetapleural carina extended on anterior 0.4 of metapleuron. Propodeum smooth and shiny with evenly sparse fine setiferous punctures on anterior half and laterally, in dorsal view 1.0–1.1× as long as medially wide. Propodeal spiracle about 0.14–0.18× the length to anterior end of pleural carina. Hind leg with femur 4.3–4.5× as long as high, 0.95–1.0× as long as tibia. Fore wing with vein cu-a opposite to Rs & M; vein 2 rs-m 0.45–0.55× as long as abscissa of M between 2 rs-m and 2 m-cu; abscissa of Cu 1 between 1 m-cu and Cu 1 a 1.55–1.6× as long as Cu 1 b. Hind wing with vein cu-a 0.35× as long as first abscissa of Cu 1 between M and cu-a, vein cu-a reclivous, first abscissa of Cu 1 vertical and straight; sometimes vein cu-a + abscissa of Cu 1 between M and cu-a straight or slightly curved, vertical; distal abscissa of Cu 1 very weakly pigmented to absent.

Metasoma ( Figs 46A, 46I–J ). Tergite I 1.55–1.65× as long as posteriorly broad, shiny, with sparse shallow setiferous punctures laterally and posteriorly, spiracle near its anterior 0.35–0.4; lateromedian longitudinal carinae present at anterior 0.1–0.2 of tergite, very weak; lateral longitudinal carinae present at posterior 0.2–0.5 of tergite. Sternite I extending posteriorly 0.6–0.7 the length of tergite. Tergite II 1.17–1.3× as long as posteriorly broad, with dense moderately deep punctures, distance between punctures about its diameter. Ovipositor very slender, slightly and evenly up-curved at 0.4 distal, 1.8–1.85× as long as hind tibia. Ovipositor sheath about 1.45×as long as hind tibia, ventral margin basally without denticles, length of setae on average 1.5–2.0× the sheath basal width.

Colouration ( Fig. 46 ). Head mostly white cream, with apex of mandibles, fovea, narrow longitudinal stripe at middle of face, frons and vertex medially, middle part of genal orbit, mid and dorsal posterior part of gena and occipital region, dark brown to black; antenna brown, scapus, pedicel and annellus and first and sometimes second flagellomere, lateroventrally white cream. Mesosoma mostly orange; dorsal rim of propleuron, mid lateral band of pronotum and dorsal part of propodeum from dark orange to black; propleuron mostly, dorsal and ventral lateral bands on pronotum, two longitudinal stripes on mesoscutum posterior to notauli, tegula, subalar prominence, scutellum dorsally, postscutellum, ventral part of mesopleuron, mesepimeron, sometimes mesosternum, posterior spot on metapleuron and lateral parts of propodeum, white cream. Metasoma dark brown; tergites I-VI with white cream posterolateral marks, tergites II-VI with black posterior band; tergite I with yellowish lateral marks; ovipositor brown; ovipositor sheaths dark brown. Legs predominantly white cream; front leg with dorsal stripe on femur, dorsal and ventral stripes on tibia, and distal part of tarsi, brown; mid leg with base of trochanter, dorsal stripe on femur, dorsal and ventral stripes on tibia, and distal part of tarsi, brown; hind leg with lateral spots on coxa, base of trochanter, base, and lateral subdistal marks on femur, basal and distal part of tibia, and distal part of tarsi, brown. Wings hyaline, pterostigma blackish brown.

Male: Unknown.

Material examined.

Type material ( 13 ♀♀). Holotype. Costa Rica: 1 ♀, Limón Prov., 7 Km SW of Bribri, 50 m, XII-1989 ( NHMUK). Non type material. Colombia: 1 ♀, Magdalena, PNN Tayrona , Pueblito , 11 o 20'N, 74 o 02'W, 225 m, 1–19-IX-2000, malaise trap, R. Henríquez ( IAvH); 1 ♂, same locality, collector and trap type, 14–29-VII-2000 ( IAvH); 1 ♀, same locality, collector and trap type, 30-IX/ 26-X-2000 ( IAvH); 2 ♀♀, same locality, collector and trap type, 26-X/ 22-XI-2000 ( IAvH); 4 ♀♀, 1 ♂, same locality, collector and trap type, 13-I/ 21-II-2001 ( IAvH). Costa Rica: 1 ♀ Cartago, Dulce Nombre, Vivero, Linda Vista, 1300m, VII-1993, Hanson, det Gauld 91 pronoto más oscuro ( NHMUK); 1 ♀, Valle La Estrella , Prov. Limon, 100m, 8-IV/ 15-V-1994, G. Carballo, Malaise, L _N_ 184600_643300 #3146 ( MNCR). Venezuela: 1 ♀, Tucuco, Zulia, 23-IV-1981, H. K. Townes ( EMUS); 1 ♀, same data, 26-IV-1981 ( EMUS).

Distribution. Colombia *, Costa Rica, Venezuela * ( Fig. 51B , Appendix I).

Figures

FIGURE 46. Clistopyga stanfordi, female (holotype): (A) habitus (lateral view); (B) head (frontal view); (C) epomia (arrow); (D) head and mesoscutum (dorsal view); (E) head and mesosoma (lateral view); (F) metapleuron and propodeum (lateral view); (G) fore wing; (H) hind wing; (I) metasoma (dorsal view); (J) ovipositor (lateral view). Scale bar: 1 mm (A, E); 0.5 mm (D, I, J); 0.2 mm (B, C, F, G, H).

FIGURE 51. Geographic distribution of C. calixtoi species group: (A) C. nasa sp. nov., C. nubiphila sp. nov.; (B) C. carvajali Gauld, Ugalde & Hanson, C. meridae sp, nov., C. stanfordi Gauld.