Clistopyga carvajali Gauld, Ugalde & Hanson, 1998

Bordera, Santiago, Palacio, Edgard & Herrera-Flórez, Andrés Fabián, 2025, The Neotropical species of Clistopyga (Hymenoptera, Ichneumonidae, Pimplinae). Part VI: the C. calixtoi species group, with the description of twenty-four new species, Zootaxa 5662 (1), pp. 1-115 : 32-35

9. Clistopyga carvajali Gauld, Ugalde & Hanson, 1998

( Figs 14 , 51B , Appendix I)

Clistopyga carvajali Gauld, Ugalde & Hanson, 1998: 167 . Holotype: ♀, Costa Rica (MNCR).

Diagnosis. Clistopyga carvajali may be distinguished from all other species of the C. calixtoi species group by the following combination of characters: mesopleuron orange, with a wide ventral longitudinal band and subalar prominence white cream ( Fig. 14F ); ventral margin of pronotum entirely white cream banded ( Fig. 14A, 14F ); metasoma mostly black with posterior well defined white cream wide bands on tergites I–V, tergites VI-VIII orange ( Figs 14A, 14G ); hypostomal carina lamelliform posteriorly to mandible base ( Fig. 14D , arrow); epomia as a strong short vertical carina ( Fig. 14F ); propodeal spiracle 0.10–0.15× the length to anterior end of pleural carina (Fig. (14F). tergite II smooth and shiny centrally ( Fig. 14G ); ovipositor 2.5–2.7× as long as hind tibia ( Fig. 14A, 14I ); ovipositor sheath basally without denticles, 2.2–2.3× as long as hind tibia; hind wing with distal abscissa of Cu1 weakly pigmented distally ( Fig. 14H ).

Description. Female: Body length 7.0– 10.3 mm. Fore wing length 4.7–7.2 mm.

Head ( Figs 14A–F ). In dorsal view, strongly narrowed behind eyes. Gena shiny, mostly smooth with very sparse setiferous punctures, in dorsal view straight, 0.26–0.3× as long as eye, in frontal view moderately constricted below eyes with straight profile. Frons and vertex smooth and shiny, with sparse setiferous punctures. Posterior ocellus separated from eye 1.0–1.1 its maximum diameter. Distance between hind ocelli 0.76–0.85× maximum diameter of posterior ocellus. Hypostomal carina lamelliform and translucent behind mandible. Face with shallow and moderately dense punctures, distance between punctures more than twice the diameter of punctures. Clypeal suture moderately curved medially. Clypeus 1.65–1.75× as broad as medially long, weakly convex with very few punctures dorsally, flat and smooth ventrally, ventral margin almost straight. Malar space about 0.8–0.84× as long as basal mandibular width, with a slightly granulate narrow subocular sulcus. Antenna with 26–28 flagellomeres, first flagellomere 6.85–7.0× as long as medially wide.

Mesosoma ( Figs 14A, 14E–F, 14H ). Pronotum smooth and shiny, almost glabrous, with fine and scattered setiferous punctures in dorsal posterior part. Epomia as a strong short vertical carina. Mesoscutum mostly smooth and shiny, with very fine and sparse shallow setiferous punctures, lateral lobes virtually glabrous; anterior half of median lobe not conspicuously prominent, confluent with posterior half. Notaulus moderately deep, reaching about 0.5 the length of mesoscutum. Mesopleuron smooth and shiny, with sparse small setiferous punctures. Epicnemial carina weak, its dorsal end straight, ending at level of ventral third of pronotum. Metapleuron smooth and shiny, glabrous, except for some sparse setiferous punctures at posterior part, 2.85–3.0× as long as high. Submetapleural carina absent. Propodeum shiny with evenly very sparse fine setiferous punctures, in dorsal view 1.45–1.50× as long as medially wide. Propodeal spiracle 0.10–0.15× the length to anterior end of pleural carina. Hind leg with femur 3.7–3.85× as long as high, 1.0–1.1× as long as tibia. Fore wing with vein cu-a slightly proximal to Rs & M; vein 2 rs-m 0.4–0.45× as long as abscissa of M between 2 rs-m and 2 m-cu; abscissa of Cu 1 between 1 m-cu and Cu 1 a 1.55–1.7× as long as Cu 1 b. Hind wing with vein cu-a 0.35–0.45 × as long as abscissa of Cu 1 between M and cu-a, vein cu-a slightly reclivous, first abscissa of Cu 1 vertical; distal abscissa of Cu 1 absent or weakly insinuated.

Metasoma ( Figs 14A, 14G, 14I ). Tergite I 1.60–1.7× as long as posteriorly broad, shiny, with very sparse shallow setiferous punctures laterally and posteriorly, spiracle near its anterior 0.35–0-4; lateromedian longitudinal carinae present at anterior 0.2 of tergite; lateral longitudinal carinae absent or slightly present at posterior 0.2 of tergite. Sternite I extending posteriorly about 0.5 the length of tergite. Tergite II smooth and shiny, 1.1–1.3× as long as posteriorly broad, with sparse shallow setiferous punctures, distance between them much more than its diameter; punctation of posterior tergites tending to be denser. Ovipositor moderately up-curved at posterior 0.3, 2.5–2.7× as long as hind tibia. Ovipositor sheath 2.2–2.3× as long as hind tibia, ventral margin basally without denticles, length of setae on average 2.5–2.8× the sheath basal width.

Colouration ( Fig. 14 ). Head mostly white cream, with apex of mandibles, lateral parts of clypeal suture, fovea, frons and vertex medially, posterior part of gena and occipital region, dark brown to black, sometimes mid part of genal orbit interrupted with black; antenna brown, scapus, ventrally whitish. Mesosoma mostly orange; propleuron dorsally, mid lateral part of pronotum, ventral part of metapleuron, lateral parts of scutellum and postscutellum and most part of propodeum dark brown to black; propleuron ventrally, dorsal and ventral wide bands on pronotum, subalar prominence, tegula, mesepisternum, ventral part of mesopleuron diffusely, two longitudinal stripes posterior to notauli, scutellum and postscutellum dorsally, dorsal part of metapleuron and widely lateral parts of propodeum, white cream. Metasoma mostly dark brown; tergite I laterally and posteriorly, anterolateral spots on tergites I–III and posterior wide band on tergites II–VI, white cream, tergites II–III(IV) with lateral posterior black spots, tergites VI–VIII orange; ovipositor dark brown; ovipositor sheaths black. Legs predominantly white cream; front leg with dorsal stripe on femur, tibia and distal part of tarsi, brown; mid leg with base of trochanter, dorsal stripe on femur, dorsal and ventral stripes on tibia, and distal part of tarsi, brown; hind leg with anterior and posterior spots on coxa, base of trochanter, distal part of trochantellus, base, and anterior and posterior subdistal marks on femur, basal and distal part of tibia, and distal part of tarsi, brown. Wings hyaline, pterostigma blackish brown.

Male: Unknown.

Material examined.

Type material. Paratypes (2 ♀♀). Costa Rica : 1 ♀, Cartago Pv. Linda Vista, Dulce Nombre , 1300m, VI/ VII-1993, P. Hanson. Det Gauld 1988 ( NHMUK) ; 1 ♀, Guanacaste Pv., Guanacaste NP, 1000m, Est. Mengo, Volcan Cacao, V-1988, Gauld & Janzen ( NHMUK) .

Non type material (4 ♂♂). Colombia : 1 ♀, Valle, PNN Farallones de Cali, Los Andes , vda Quebradahonda , 3 º 34'N, 76 º 40'W, 1730 m, 20-V-1998, net trap, W.Alfonso ( MJHN) . Costa Rica: 1 ♀, Prov. Guanacaste, La Cruz, P.N. Guanacaste, Estación Pitilla , 9km S. de Santa Cecilia, 700m, II-1995, P. Rios, Malaise Trap, L_N_329950_380450, #4517 ( MNCR) ; 1 ♀, San Vito de C. B. las Cruces , 1200m, 17-VIII/ 12-IX-1982. B. Gill. ( EMUS) ; 1 ♀, Sendero Nacho, Est. Pitilla, Prov. Guanacaste, 700m, V-1994, P. Rios, Malaise, L _N 330200_380200, #3339 ( MNCR) .

Distribution. Colombia *, Costa Rica ( Fig. 51B , Appendix I).

Figures

FIGURE 14. Clistopyga carvajali, female: (A) habitus (lateral view, paratype); (B) head (frontal view, holotype); (C) head (dorsal view); (D) head (lateral view, lamelliform hypostomal carina, arrow); (E) head and mesosoma (dorsal view); (F) head and mesosoma (lateral view); (G) metasoma (dorsal view); (H) wings; (I) ovipositor (lateral view). Scale bar: 1 mm (A, G); 0.5 mm (E, F, H, I); 0.2 mm (B, C, D).

FIGURE 51. Geographic distribution of C. calixtoi species group: (A) C. nasa sp. nov., C. nubiphila sp. nov.; (B) C. carvajali Gauld, Ugalde & Hanson, C. meridae sp, nov., C. stanfordi Gauld.