Bryopharsos Quate, 1996

Jaume-Schinkel, Santiago, Kilian, Isabel C., Pazmiño-Palomino, Alex & Mengual, Ximo, 2025, Revision of the genus Bryopharsos Quate, 1996 (Diptera: Psychodidae) with the description of nine new species, European Journal of Taxonomy 1001 (1), pp. 1-51 : 4

Genus Bryopharsos Quate, 1996

Bryopharsos Quate, 1996: 40 . Type species: Bryopharsos palpiculum Quate, 1996 by original designation.

Bryopharsos – Quate 1999: 434 (female description and updated distribution). — Ježek 2010: 237 (discussion of characters). — Kvifte 2018: 603 (tribal classification). — Bravo & Araújo 2019: 364 (description of new species and key to world species).

Species distribution shown in Fig. 1 .

Diagnosis

Building upon Quate’s (1996) original diagnosis, Bravo & Araújo (2019) proposed a broader diagnosis to accommodate their new Brazilian species. In this study, we update the genus diagnosis, including previously and herein described species. Male: vertex short, up to twice as wide as eye bridge; eye bridge contiguous conformed by 3–5 facet rows; antennae with 14 flagellomeres, 1–13 asymmetricallynodiform, flagellomere 14 with long, digitiform apiculus; flagellomeres 1–13 with a pair of broad, leafshaped ascoids; ascoids width at least the same as the length of the flagellomere carrying them; palpi short reaching the level of the third flagellomere; labellum short and not bulbous; thorax with or without allurement organs; wing membrane with alveoli between longitudinal veins; wing vein R 5 ends at wing apex; gonocoxal apodemes fused or contiguous at midline; gonocoxal lobes projected anteriorly, with or without a spine; gonostyli digitiform to club-shaped with rounded apex; surstyli with 1–7 tenacula; aedeagal complex asymmetrical; ejaculatory apodeme width is equal to or greater than half of its total length in ventral view.

Bryopharsos was considered by Ježek (2010) to be morphologically close to the Palearctic/Oriental genus Saximormia as they share the following characters: ascoids as broad as or broader than the flagellomere carrying them, eyes contiguous in the eye bridge, and the general wing structure of the wing. Nevertheless, they can be easily differentiated from each other by the following characters: alveoli on the wing membrane present in Bryopharsos (absent in Saximormia ), wing vein radial and medial forks being at the same level and basal to the wing center in Bryopharsos (forks not being at the same level and apical to the wing center in Saximormia ), vein R 5 ending at wing apex in Bryopharsos (vein R 5 ending at wing margin posterior to the apex in Saximormia ), the ejaculatory apodeme is conspicuously enlarged in ventral view in Bryopharsos (ejaculatory apodeme not enlarged in Saximormia ), gonostyli club-shaped with a rounded blunt apex in Bryopharsos (gonostyli tapering towards the apex and with a pointed apex in Saximormia ) ( Ježek 2010; Bravo & Araújo 2019).

Figures

Fig. 1. Species distribution map. Species dots do not represent the actual localities; jitter was added for visualization as the species’ real distributions overlap.