Spodochlamys, Burmeister, 1855

Kobayashi, Kazuo & Ratcliffe, Brett C., 2023, A New Species of Spodochlamys Burmeister, 1855 (Coleoptera: Scarabaeidae: Rutelinae: Anatistini) from Colombia and a Key and Checklist of Spodochlamys Species, The Coleopterists Bulletin 77 (1), pp. 120-126 : 124-125

KEY TO SPECIES OF MALE SPODOCHLAMYS

Note: Males of Spodochlamys letiranti Jameson and Ratcliffe are not known. The four Soula species ( S. peruvianus , S. lydiae , S. curoei , S. jamesonae ) are not included in the key for lack of known characters. See Diagnosis in text.

1. Head and pronotum brown, black, or cupreous, occasionally with metallic green reflections (with magnification and illumination). Elytral epipleuron (ventral view) lacking short, dense setae. Elytral surface lacking minute setae .... 2

1′. Head and pronotum greenish in color (with magnification and illumination). Elytral epipleuron (ventral view) with short, dense setae. Elytral surface with minute setae ..... 13

2. Pronotum distinctly vaulted in lateral view. Protibia greatly elongate and bowed. Elytra with waxy bloom ............. S. iheringi Ohaus

2′. Pronotum regularly convex in lateral view. Protibia simple, not elongate and not bowed. Elytra with or without bloom........................ 3

3. Clypeal apex strongly narrowed to a recurved, acute, narrowly bidentate tooth. Elytra with waxy bloom.............. S. caesarea Burmeister

3′. Clypeal apex rounded, lacking strong attenuation. Elytra without waxy bloom ................ 4

4. Pygidial disc protuberant in lateral view ..... 5

4′. Pygidial disc evenly convex in lateral view, not noticeably produced posteriorly.................... 9

5. Inner claws (all legs) simple. Apical umbone of elytra rounded, lacking spinose projection ................................ S. flavofemorata Ohaus

5′. Inner claw of meso- and metathoracic legs split, prothoracic leg with claw weakly split or simple. Apical umbone of elytra with spinose projection or tubercle ......................................... 6

6. Posterior margin of metafemur not narrowly laminate, not produced posteriorly ............... 7

6′. Posterior margin of metafemur narrowly laminate and produced posteriorly ............... 8

7. Inner metatibial spur strongly hooked. ........... ...... S. ancistrospina Jameson and Ratcliffe

7′. Inner metatibial spur not hooked ................... .............................................. S. soulai Curoe

8. Apical umbone of elytra with well developed, spinose projection ................. S. feyeri Ohaus

8′. Apical umbone of elytra tuberculate .............. ................... S. neitai Jameson and Ratcliffe

9. Apex of labrum acutely angulate ................... ............................................ S. gigas Murray

9′. Apex of labrum obtusely angulate .............. 10

10. Inner claw split at apex on all legs ................. ........................................... S. cupreola Bates

10′. Inner claw of prothoracic leg simple; claw split or not on meso- and metathoracic legs. ............................................................ 11

11. Elytral color dark brown, opaque ................... .......... S. marahuaca Jameson and Ratcliffe

11′. Elytral color reddish brown, shiny or with waxy bloom .......................................................... 12

12. Elytra with waxy bloom.................................. ................ S. neblina Jameson and Ratcliffe

12′. Elytra shiny, with weak metallic copper reflection ................ S. keitai Kobayashi and Ratcliffe, new species

13. Pronotum greatly vaulted in lateral view; in dorsal view, basolateral margins greatly explanate. Anterior margin of profemur with well developed, forward-projecting spine near apex............................................................. 14

13′. Pronotum regularly convex in lateral view; in dorsal view, basolateral margins not greatly explanate.Anterior margin of profemur lacking forward-projecting spine near apex ............ 15

14. Anterior margin of elytra with weak, external tooth. Mesotibia at apex acutely produced on external edge. Metatibia not greatly flattened in lateral view ........ S. mirabilis Waterhouse

14′. Anterior margin of elytra without external tooth. Mesotibia at apex quadrate, not produced on external edge. Metatibia greatly flattened in lateral view .......................... S. latipes Arrow

15. Pronotum on lateral margin near middle rounded, lacking sharp angulation. Pronotal surface rugopunctate ................... S. nazareti Arnaud

15′. Pronotum on lateral margin near middle with sharp angle. Pronotal surface densely punctate....................................................... 16

16. Inner margin of protibia weakly curved. Mentum with median, longitudinal, parallel-sided depression that extends from apex to base........... S. curvibrachialis Ohaus

16′. Inner margin of protibia straight, not weakly curved. Mentum with median depression that extends from apex to mid-disc ....................... ........................................... S. poultoni Shipp