Thyanta (Thyanta) perditor (Fabricius)

Rider, David A. & Chapin, J. B., 1992, Revision of the genus Thyanta Stal, 1862 (Heteroptera: Pentatomidae) II. North America, Central America, and the West Indies., Journal N Y Ent Soc 100 (1), No. 1, pp. 42-98 : 64-69

Thyanta (Thyanta) perditor (Fabricius)

Figs. 63-77, Map 5

Cimex perditor Fabricius, 1794:102 ; Fabricius, 1803:163.

Pentatomafascifera Palisot de Beauvois, 1817: 150, fig. 8 (syn. by Dallas, 1851: 206). Pentatoma collaris Westwood, 1837:40 (syn. by Dallas, 1851: 206).

Cimex transversalis Herrich-Schaffer, 1841:66 (syn. by Dallas, 1851:206).

Cimex dimidiatus Herrich-Schaffer, 1841 :fig. 629 (syn. by Dallas, 1851: 206).

Pentatoma dimidiatum: Herrich-Schaffer, 1844:94 (syn. by Dallas, 1851:206).

Euschistus perditor: Dallas, 1851:206 ; Walker, 1867:247.

Pentatoma (Mormidea) perditor: Guerin-Meneville, 1857:367 .

Thyantaperditor: Stal, 1862a:58; Stal, 1862b: 104; Stal, 1868:29; Stal, 1872:34; Uhler, 1872: 399 (part); Uhler, 1876:289; Uhler, 1877:404 (part); Distant, 1880:66; Berg, 1884: 100; Distant, 1893:333; Lethierry and Severin, 1893: 148; Uhler, 1893:705; Uhler, 1894a:230 (part); Uhler, 1894b: 173; Distant, 1900:432; Van Duzee, 1904: 52-53 (part); Van Duzee, 1907:9; Kirkaldy, 1909:95; Banks, 1910:90; Zimmer, 1912: 14 (part); Barber, 1914: 523; Van Duzee, 1917:51-52; Barber, 1923:12; Blatchley, 1926: 113, 114-115 (part); Barber, 1939:292-293; Torre-Bueno, 1939: 230; Ruckes, 1957a:1, 20; Froeschner, 1988:593.

Euschistusfasciatus Walker, 1867:245 (syn. by Stal, 1872:34).

Euschistus adjunctor Walker, 1867:249 (syn. by Stal, 1872:34).

Thyanta (Thyanta) perditor : Rider and Chapin, 1991.

Diagnosis. Transhumeral rubiginous band usually present; tylus and vertex ofhead often reddish.

Lateral jugal margins sinuous, not parallel ( Fig. 64). Humeral angles spinose, each spine directed anterolaterad; anterolateral pronotal margins not piceous, concave in dorsal view ( Fig. 63). Mesial comer of each pronotal cicatrice black. Each abdominal stemite with postspiracular black spot on each side, both anterolateral and posterolateral angles usually piceous.

Basal plates with mesial margins straight to slightly convex in caudoventral view; posterior margins sinuous ( Fig. 75). Pygophoral opening subtended by semicircular impression; posterior margin of pygophore produced posterodorsad, in ventral and dorsal views convex medially with small, medial, V-shaped emargination ( Figs. 72, 73), concave in lateral view ( Fig. 74).

Types. Fabricius (1794) described Cimex perditor from 25<3 and 2'i''i' specimens without designating a holotype or paratypes. Rider and Chapin (1991) made lectotype and paralectotype designations. All four specimens, which are housed in the Universetetes Zoologiske Museum (Copenhagen, Denmark), were examined.

Rider and Chapin (1991) confirmed the status of Pentatoma fascifera Palisot de Beauvois, P. col/aris Westwood, Cimex transversalis Herrich-Schiiffer , Euschistus fasciatus Walker , and E. adjunctorWalker as junior synonyms. They also designated lectotypes for the latter two species. At one time Euschistus rubiginosus Dallas was considered a synonym of T. perditor. Rider (1986b), however, examined the holotype of E. rubiginosus and determined that it was a senior synonym of Euschistus incus Rolston.

Distribution. This is the most widely distributed species in the genus, occurring from southern Florida, Texas, and Arizona south through Central America, West Indies, and South America to northern Argentina ( Map 5).

Specimens examined. 844 specimens collected during every month of the year; deposited in AMNH, AUA, BMNH, CAS, CNC, CU, DAR, DBT, FSCA, INHS, ISU, LHR, LSU, MSU, MSUE, OSUC, PUL, SIUC, SMEK, TAMU, UAT, UCB, UCR, UCS, UGA, UMAA, UMC, UNAM, USNM, WSU. UNITED STATES: Arizona: Cochise: Southwest Research Station , 5 mi W Portal . Florida: Flamingo ; Mahogany Hammock ; Paradise Key ; Pine Castle ; Windly . Broward: Deerfield Beach . Collier: Immokalee ; Royal Palm Park . Dade: Coral Gables ; Everglades National Park ; Florida City ; 5 mi SW Florida City ; Goulds ; Grossman Hammock ; Hialeah ; Homestead ; Kendall ; Miami ; Miami Springs ; Princeton . Highlands: Avon Park ; Lake Placid ; Sebring . Indian River: Vero Beach . Lee: Fort Myers . Manatee: Palmetto ; Terra Ceia . Martin: Stuart . Monroe: Big Pine Key ; Key Largo ; Key West ; Marathon Key ; Upper Key Largo ; Upper Matecumbe Key . Orange: Orlando . Palm Beach: Belle Glade ; Delray ; Royal Palm Park . Polk: Lake Hamilton ; Lakeland . Volusia : Daytona Beach . Texas: Cameron: Brownsville ; Sabal Palm Grove near Southmost .

MEXICO: Hochmilco ; Lagos de Moreno ; Paricutin . Aguascalientes: Aguascalientes . Campeche: km 71 Carr . Campeche-Meizina . Chiapas: Arriaga ; 4.4 mi N Bochil ; Bonampak Ruinas ; Chicoasen ; Chorreadera State Park ; Cintalpa ; 5 mi S Cintalpa ; 13 mi W Cintalpa ; Comitan ; 31.5 mi SE Comitan ; Dos Lagos ; El Sumidero ; 1.6 mi S Jitotol ; 13 mi S La Trinitario ; 18 km S La Trinitario ; 12 mi W Ocozocoautla ; Palenque ; Palenque Ruinas ; 23 mi S Palenque , 4 mi NE Pichucalco ; 1 mi S Rayon ; 2 mi SE Revolucion Mexicana ; 3 mi W Rizo de Oro ; Sanches Ranch Las Rosas ; San Cristobal de las Casas ; 8 mi SE San Cristobal las Casas ; 23 mi W San Cristobal de las Casas ; 39 mi E San Cristobal las Casas ; Simojovel ; 10 km WNW Soyalo ; 2 mi E Suchiapa ; Sumidero Canyon ; 9 mi SE Tapilula ; Teopisca ; 14 mi N Tuxtla Gutierrez ; 3.5 km S Rio Tulija . Colima: Colima Volcano . Durango: 9 mi W La Ciudad ; Peasa Pena . Guanajuato: Leon . Guerrero: Acahuizotla ; Acapulco de Juarez ; 5 mi S, 2.5 mi E Chilpancingo ; km 8 Carr . Chilpancingo Omilteme ; 17 mi N Mexcala ; Mochitlan ; 1.5 mi W Mochitlan ; 13 mi SW Tierra Colorado . Hidalgo: Hwy 45 , 17 mi NE Huichapan . Jalisco: 16 km E Agua el Obispo ; Ajijic ; Guadalajara ; 5 mi SE Plan de Barrancas . Mexico: 4.3 mi NE Ixtapan de la Sal ; Tejupilco ; Temascaltepec , Real de Arriba ; Teotihuacan ; 11 mi W Texcoco ; 1 mi NE Tlamacas , P. N. Popocatepetl ; Valle de Bravo ; 21 mi NW Valle de Bravo . Michoacan: Apatzingan ; 11 mi E Apatzingan ; Jiquilpan ; Palo Alto ; Tancitaro . Morelos: Cuautla ; Cuernavaca ; Jojutla ; Morelos ; Pte de Itla ; Xochicallo . Nayarit: 13 mi NW Ahuacatlan ; Jesus Maria ; NayaritJalisco line , Hwy 15 ; 15 km E San Blas ; Tepic . Nuevo Leon: 9 mi S Monterrey ; 3 mi S Pacheco . Oaxaca: 7.7 mi S Ejutla ; El Camaron , 20 mi E Oaxaca ; 2.7 mi NW El Camaron ; 10 mi S El Camaron ; 27 mi E Juchitan ; 3 km E La Ventosa ; 3.4 mi SE Matatlan ; 11 mi N Matias Romero ; SE Nejapa ; 30 mi NW Oaxaca ; 45 mi SE Oaxaca ; 53 mi NE Oaxaca ; Tehuantepec ; Temazcal ; 23 mi W Tequistlan ; Totolapan ; 18 mi NW Totolapan ; 23 km NW Totolapan ; Tuxtepec . Puebla: 5.1 mi SW Tehuacan ; 6 mi SW Tehuacan . Queretaro: Queretaro ; 29 mi N Queretaro ; 10 mi E San Juan del Rio . San Luis Potosf: 5 mi E Ciudad del Maiz ; 11 km N Ciudad Valles ; 20 mi S Ciudad Valles ; El Pujal ; El Saito Falls ; Micos ; 31 mi S San Luis Potosi ; Tamazunchale ; 30 mi S Tamazunchale ; Valle Hidalgo ; 2 mi E Xilita . Sinaloa: 22.6 mi S Culiacan ; Los Mochis ; Mazatlan ; 15 mi N Mazatlan ; Valle Union ; 5 mi E Valle Union . Tabasco: 30 mi W Cardenas ; Chontalpa ; Pajelagatero ; Pico do Oro ; 5 km S Villahermosa ; 14 mi SE Villahermosa . Tamaulipas: Bocatoma: 11 mi SW Ciudad Victoria ; 6 mi W Rio Sabinas , near Encino ; 19 mi NE Tula . Tlaxcala: 2 mi W Tlaxcala . Veracruz: Alvarado ; Catemaco ; 7 mi SE Catemaco ; Coatzacoalcos ; Cordoba ; 1 mi E Cordoba ; Cotaxtla ; Extacion Mieron das Fortin ; 1 mi W Fortin de las Flores ; Jalapa ; 10 mi E Jalapa ; Lake Catemaco ; 5 mi N Lerdo de Tejada ; Los Tuxtlas Biological Station ; L. Rivera . Atzagan ; 4 mi NE Minatitlim ; Nanchital ; Orizaba ; Papantla ; Plan del Rio ; 3 mi SW Sontecompapan ; 15.8 mi S Tampico ; Vega de Alatorre ; Veracruz . Yucatan: 10 km N Piste .

GUATEMALA: Chocola . Chimaltenango: Yepocapa . Guatemala: Guatemala City . Izabal: Livingston ; Los Amates ; Morales . Jutiapa: San Jeronimo . Sac.: Antigua Guatemala . Solo/a: Panajachel , Lago de Atitlan . Suchitepequez: Alta Vista ; 8 mi N Santa Barbara . BELIZE: Punta Gorda . EL SALVADOR: La Libertad ; San Salvador . HONDURAS: 1 mi W Jicaro Galan Junction , Rio Nacaome ; Tegucigalpa ; Uyace Peak ; Yojoa Lake ; Zomorano . NICARAGUA: km 4 to Masaya . Estelf: 13.4 mi NW Sebaco . Leon: Los Zarzales ; Malpaisillo . Managua: Los Brasiles ; Managua ; Tipitapa . Musawas: Waspuk R . Zelaya: Puerto Cabezas . COSTA RICA: Isla Bonita ; Reventazon Valley ; San Carlos . Cartago: Santiago ; Turrialba . Guanacaste: La Pacifica near Canas . Heredia: Heredia ; 2 mi E Puerto Viejo ; Varablanca . Puntarenas: Monteverde . San Jose: 5.5 km SE Desamparados ; San Jose . PANAMA: New California . Bocas de! Toro: Chiriqui Grande . Canal Zone: Barro Colorado Island ; Cristobal . Chiriquf: Boquete ; Cerro Punta ; Porteillos . Cocle: Cerro Pena near El Valle . Panama: Cerro Jefe ; Madden Lake .

BAHAMA ISLANDS: Andros Island: Andros Town . Cat Island . Long Island: Clarencetown . New Providence Island: Nassau . CUBA: El Cobre . Ciudad de la Habana: La Habana . Cienfuegos: Palmira ; Soledad nr Cienfuegos . Granma: Cayamas . Guantanamo: Mtns nr Guantanamo . Las Tunas: Jobabo . Pinar def Rio: San Vicente . Santiago de Cuba: 12 mi N Santiago de Cuba . JAMAICA: Balaclava ; Claremont Baron Hill Trelawny ; Hope Bay ; Kingston ; Liguanea Plain ; Mandeville ; Montego Bay . HAITI: Ouest: Kenscoff ; Port-au-Prince . DOMINICAN REPUBLIC: 8 mi up Macoris River . Distrito Nacional: Santo Domingo . La Romana: La Romana Center . La Vega: Jarabacoa . Peravia: 2 km N Nizao ; 21 km NW San Jose . Puerto Plata . San Cristobal: San Cristobal . San Pedro de Macorfs: Boca Chica . PUERTO RICO: Cialitos Cruces , El Verde , Mona Island . Aguadilla: Aiiasco , lsabela , Rincon , San Sebastian . Arecibo: 7 km S Ciales , Dorado , Manati , Utuado , Vega Baja . Bayamon: Corozal . Guayama: Arroyo . Humacao: Humacao , Loiza , 1 mi SE Luquillo , Naguabo . Mayaguez: Boqueron , Guanica Forest , Guayanilla , Mayagiiez , San German . Ponce: Coamo Springs , Juana Diaz , Ponce , 3 mi N Santa Isabel . San Juan: Rio Piedras . U. S. VIRGIN ISLANDS: St. Croix: Frederiksted . St. Thomas . BRITISH WEST INDIES: Antigua: Bello . Barbuda: Codrington . Dominica: Castle Bruce Road nr Savane David ; Clarke Hall Est ; 4. 7 mi E Pt Casse ; 5 km SW Pt Casse ; Roseau . Grand Cayman: 3 mi N Georgetown ; Western Dist . Grenada: Grand Anse ; Mount Gay Est ; St. Georges . St. Kitts: Basseterre ; 4 mi W Basseterre . St. Lucia: Castries . St. Vincent . Trinidad: Cumuto ; St. Augustine . FRENCH WEST INDIES: Guadeloupe: Pointe-a-Pitre . Martinique: 2 km N St. Pierre . BARBADOS .

Comments. Thyanta perditor is one of a group of very closely related species, all of which have distinctly spinose humeral angles. The characters used to separate these species are sometimes subtle and hard to diagnose unless a series of specimens is available. Fortunately, most have very little overlap in their distributional ranges.