Haploceras staszycii ( Zejszner, 1846 )

Vašíček, Zdeněk & Skupien, Petr, 2025, Revision of haploceratid ammonoids from the Štramberk Limestone, Jurassic-Cretaceous boundary beds (Outer Western Carpathians), Acta Palaeontologica Polonica 70 (3), pp. 421-441 : 427-431

Haploceras staszycii ( Zejszner, 1846)

Figs. 3B , 4B , 8A–D.

1846 Ammonites Staszycii Zejszner 1846 : pl. 4: 3a–c.

1870 Haploceras Staszycii Zeuschner ; Zittel 1870: 50, pl. 27: 2a, b, 3a, b (neotype), non figs. 4, 5 (?).

1876 Haploceras Staszycii Zeuschn. ; Gemmellaro 1876: 34, pl. 7: 1–3.

1879 Haploceras staszycii Zeuschner ; Fontannes 1879: 11, pl. 2: 4a, b.

1960 Haploceras staszycii Zeuschn. ; Collignon 1960: pl. 142: 539.

→ Fig. 6. Haploceratid ammonoids from Tithonian–lower Berriasian, Upper Jurassic–Lower Cretaceous. A, B. Hypolissoceras woehleri ( Oppel, 1865) . A. Holotype AS III 811 (real state of preservation), external mould, Štramberk, in lateral (A 1) and ventral (A 2) views. B. Z4171, external mould, Štramberk, in lateral (B 1) and ventral (B 2) views. C. Volanites verruciferus ( Zittel, 1869) , Z2981b, external mould, Štramberk, in lateral (C 1) and ventral (C 2) views. D. Hypolissoceras subtilius ( Zittel, 1870), Z2767a, external mould, Štramberk, in lateral (D 1) and ventral (D 2) views, view of the area of peristome (D 3). E. Beyrichilamellaptychus beyrichi beyrichi ( Oppel, 1865) , Ge0001, lateral view, from Měchová et al. (2010: fig. 9B), Kurovice. F, G. Haploceras elimatum ( Oppel, 1865) . F. Specimen with Punctaptychus punctatus ( Zittel, 1868) in the body chamber, AS III 339, in lateral view, from Barthel (1962), according to Barthel’s hand-written label: “ H. elimatum (Oppel) mit erhaltenen Aptychen, Tithon, Le Pouzin, Typloc. zu Toucas 1890 ”. G. Z1824, specimens car- rying the calcitic shells of serpulid worms (white dots), Štramberk. The straight line perpendicular to periphery of specimen shows the margin between body chamber and phragmocone. Photos Kristýna Mezihoráková (Ostrava, Czechia), exept A, ZV; and G. Ondřej Malek, (Green Gas DPB, Paskov, Czechia).

Material. — The material under this study consists of 8 specimens from Moravian-Silesian museums (e.g., Z1585, Z1484, MM 010, MM Ge 30259, B13570 , PL2447, PL5059, and PL5061 ). The set was supplemented by direct study of origi- nals in the Zittel collection housed in Munich ( SNSB-BSPG AS III 803, 805, 806). The initial half of the ultimate whorl of the specimen PL5059 belongs, according to the visible suture line, to the phragmocone, and the final half to the body chamber .

Description. —Semi-involute specimens, with a high, relatively broad whorl and a narrow umbilicus. The low umbilical wall declines obliquely to the line of coiling. It continuously passes through a rounded zone to the slightly arched flanks of the whorl. The flanks also pass through a rounded zone to the medium-broad venter. On the juvenile specimen PL2447 and further on the adult specimen Z1484, a siphonal furrow is preserved on the phragmocone. The shell is

→ Fig. 8. Haploceratid ammonoid Haploceras staszycii ( Zejszner, 1846) from Štramberk, Tithonian–lower Berriasian, Upper Jurassic–Lower Cretaceous. A. PL5059, lateral view (A 1), ventral view demonstrating the ribbing on external mould (A 2), Kotouč Quarry, debris on the 9th level. The straight line perpendicular to periphery of specimen show the margin between body chamber and phragmocone. B. Z1585, lateral (B 1) and ventral (B 2) views. C. PL2447, specimen with the siphonal groove, ventral (C 1) and lateral (C 2) views. D. MM010, lateral view. E, F. Haploceras tithonium ( Oppel, 1865) . E. PL5113, internal mould with incomplete outer suture line, lateral (E 1) and ventral (E 2) views, Kotouč Quarry, site No. 4 on the 8th level ( Vašíček and Skupien 2016). F. PL5115, juvenile specimen, lateral view, Kotouč Quarry. G–J. Hypolissoceras carachtheis ( Zejszner, 1846) . G. Z1385 internal mould, lateral view (G 1) with rest of peristome by umbilicus, ventral view (G 2). H. B12689, external mould in lateral view. I. Z2981a external mould in lateral (H 1) and ventral (H 2) views. J. SNSB-BSPG AS III 47, lateral view of the lectotype (real state of preservation), Koňákov. K. Hypolissoceras woehleri ( Oppel, 1865) , MM06, external mould, lateral (K 1) and ventral (K 2) views. The straight lines perpendicular to periphery of specimen show the margin between body chamber and phragmocone. Photos Kristýna Mezihoráková (Ostrava, Czechia); except J, Alexander Nützel (SNSB-BSPG).

smooth at the beginning. On the body chambers in the specimens under study or where the original shell is peeled off to expose the steinkern, faint, blunt to indistinct, and relatively closely spaced ribs are visible.

Measurements. —The 16 measurable specimens range from 29 to 83 mm in diameter (SOM 3). The Dmax of PL5059 is 44.0 mm; the end of the phragmocone is present at a D of approximately 32.0 mm. According to our measurement from a copy of the illustration of the lost Zejszner’s holotype (1846: pl. 4: 3a, b), at Dmax = 56.5 mm, H = 27.2 (0.48), U = 10.0 (0.18), B = 21.7 (0.38). Neotype SNSB-BSPG AS III 805 at D = 73.6 mm shows H/D = 0.51, U/D = 0.19, B/D = 0.36. Values of H/D in the material under study range from 0.47–0.52, U/D from 0.17–0.22, B/D from 0.36–0.41. The regression lines H/D, U/D, B/D of Ha. staszycii are accompanied by the microconch lines of Ha. elimatum in the graphs ( Fig. 9A–C). The only clear difference in the course of the two regression lines is evident for the U/D values.

Remarks.— Malinowska et al. (1980: 444) stated that the holotype of Ha. staszycii disappeared and established the lectotype (but actually neotype). Haploceras staszycii reaches a medium size. Characteristic features include a relatively narrow umbilicus. Ribbing, if preserved, is usually weakly developed with regard to the smaller sizes of specimens of Ha. staszycii . Haploceras tithonium ( Oppel, 1865) is a species that reaches a minor size of specimens and has higher whorls.

Stratigraphic and geographic range.— Enay and Cecca (1986) state that the stratigraphic range is Hybonotum to Ponti zones. Zeiss (2001) mentions the Tithonian of the Tethyan area with emphasis on the upper Tithonian. With regard to the assumption that Ha. staszycii forms a dimorphic pair with Ha. elimatum , the stratigraphic range should be identical to that of Ha. elimatum , i.e., the Tithonian to lower Berriasian. This argument is supported by specimens in Kvantaliani (1999) from Crimea, determined, however, as Ha. elimatum . Haploceras staszycii occurs in many localities of the Mediterranean and sub-Mediterranean areas. The neotype comes from Rogoźnik in Poland (lower Tithonian of the Pieniny Klippen Zone); it is also known from France, Italy, Romania, Hungary, Crimea, and Madagascar.

Our own collections from Kotouč Quarry come from Level 8 from locality 4 ( Vašíček and Skupien 2016) with accompanying ammonites from the Fallauxi and Microcanthum ammonite zones and from the debris on Level 9.

Figures

Fig. 3. Whorl sections of haploceratid ammonoids from Štramberk, Czech Republic, Tithonian–lower Berriasian. A. Haploceras elimatum (Oppel, 1865), SNSB-BSPG AS III 800 (Zittel 1868: pl. 13: 1a) at H = 33 mm. B. Haploceras staszycii(Zejszner,1846),SNSB-BSPGASIII 805 (Zittel 1870: pl. 27: 3a) at H = 37 mm. C. Haploceras tithonium (Oppel, 1865), SNSB- BSPG AS III 798 (Zittel 1868: pl. 14: 2a) at H = 25 mm. D. Hypolissoceras carachtheis (Zejszner, 1846), SNSB-BSPG AS III 47 (Zittel 1868: pl. 15: 1c) at H = 17 mm. E. Hypolissoceras woehleri (Oppel, 1865), MM 06 at H = 27 mm.

Fig. 4. Suture lines of haploceratid ammonoids from Štramberk, Czech Republic, Tithonian–lower Berriasian. A. Haploceras elimatum (Oppel, 1865).Almostcomplete outersuture line ofZ4224at H = 11.5mm.B. Haploceras staszycii (Zejszner, 1846). Substantial part of outer suture line of PL5061 (6th level, middle) at H = 14 mm. C. Haploceras tithonium (Oppel, 1865). Outer and adventive lobes with accompanying saddles of PL5113 at H = 15 mm. D. Hypolissoceras woehleri (Oppel,1865).Adventive lobe with parts of accompanying saddles of MM08 at H = 15 mm.A, adventive lobe; E, external lobe; U2, umbilical lobe.