Clivina circumopaca, Balkenohl, 2025

Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), Zootaxa 5592 (1), pp. 1-70 : 42-43

publication ID	https://doi.org/10.11646/zootaxa.5592.1.1
publication LSID	lsid:zoobank.org:pub:0E478171-152D-4A6F-9CB5-D9C9B5ABB744
DOI	https://doi.org/10.5281/zenodo.14953742
persistent identifier	https://treatment.plazi.org/id/03F487F9-733C-FF85-53D4-A656FA64F834
treatment provided by	Plazi
scientific name	Clivina circumopaca
status	sp. nov.

Treatment

Clivina circumopaca sp. nov.

( Figs 26 View FIGURES 15–26 , 61 View FIGURES 53–67 , 94 View FIGURES 85–96 , 120 View FIGURES 110–128 )

Type material. Holotype: ♂, with labels and data: white, black printed “C Sulawesi, 50m, 1999 6km E Tambarana, 120.28,06E 1.11,15S 9.-11. July Bolm lgt.” / “ Clivina spec. helferi-group det. Balkenohl 2001 ” ( SMNS).

Paratypes: 2 ♂♂, 2 ♀♀, same data as holotype ( SMNS, CMBB) .

Description. Measurements. Body length 5.21–5.38 mm (ẍ = 5.29 mm *), width 1.54–1.58 mm (ẍ = 1.56 mm *), ratio L/W of pronotum 0.89–0.90 (ẍ = 0.89*), ratio L/W of elytra 1.79–1.89 (ẍ = 1.85*); (*n = 5). Colour glossy, fuscous. Supraantennal plate and wing of clypeus slightly translucent. Antennae, mouthparts, intermediate and hind legs yellowish.

Head about a quarter narrower than pronotum. Clypeus with central part straight, with indistinctly projecting rounded teeth laterally, separated from clypeal-wing by slight notch; wing wider than teeth of clypeus, rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled notch, supraantennal plate with margin laterally convex, more distinct at middle, clypeus, wing, and supraantennal plate indistinctly reflexed margined. Supraantennal plate extended posteriorly into an obtuse keel, separated from wing by narrow furrow. Clypeus transverse, moderately convex, with scattered small punctures, clypeal-furrow distinct. Frons slightly to moderately convex, with indistinct flat pore at middle, with scattered small punctures, separated from supraantennal plate and supraorbital carina by distinct furrow, with clypeal seta in front; supraorbital carinae moderately developed, with two supraorbital setigerous punctures at middle and end of eye level. Neck constriction closed, formed by a slight step from frons and punctures posteriorly. Eye prominent, gena small but distinct. Antenna elongate, reaching over base of pronotum. Labrum straight anteriorly, finely isodiametrically reticulated, six-setose. Mentum slightly translucent, slightly reflexed margined, with isodiametric reticulation, lobe flat, oval, with apex rectangular, median tooth higher than lobe, hollowed out at tip, clefts relatively small. Pronotum ( Fig. 61 View FIGURES 53–67 ) with disk slightly convex in lateral view, moderately convex in frontal view. Shape trapezoid to cordate, wider than long. Reflexed lateral margin smooth, nearly straight at middle, widest at posterior third; anterior angle distinct, posterior angle distinct, with distinct laterally projecting tooth; lateral channel moderately narrow, indistinctly reticulated, continuing up to base as sharp, slightly convex margin. Median line complete, engraved at middle, joining anterior transverse line. Anterior transverse line distinct, complete. Disk completely covered with micro-punctures, with few slight transverse wrinkles laterally, with isodiametric reticulation at base, bilaterally with two groups of punctures and impression in basal third, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 30°. Basal channel on flange regularly developed, somewhat rugose, with sharp carina.

Elytron ( Fig. 26 View FIGURES 15–26 ) with disk flattened in anterior half in lateral view, moderately and regularly convex in frontal view. Shape of elytra elongate, somewhat widened, less than about two-fifth longer than wide, with lateral margin distinctly convex at middle and slightly diverging, maximum width posterior middle. Humerus rounded but seemingly angled. Reflexed lateral margin indistinctly crenulate behind humerus (100-fold). Lateral channel slightly narrower at middle. Row of setigerous punctures in the lateral channel nearly regular in size and distance, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity falling convex to pedunculus, with isodiametric reticulation. Striae moderately deep, striate-punctate at middle, punctures less distinct laterally, striae one to four free at base, five and six joining at humerus, one to four ending free at apex, five and six joining apically. Intervals moderately convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven with very short obtuse step at apex, eight carinate at apex. Surface of interval one to five at middle of disk smooth and glossy, all other parts with distinct isodiametric reticulation, apically third of elytron reticulated as well as base.

Hind wing fully developed.

Lower surface. Proepisternum isodiametrically reticulated, with few fine wrinkles laterally, proepisternum with distinct narrow furrow. Sternite of abdomen with shingle-like reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with moderately wide projection posteriorly at middle where it is slightly hollowed out, with narrow marginal furrow at middle (in both sexes).

Legs. Protibia moderately slender, with fine longitudinal reticulation, with slight sulcus dorsally, with three slender acute spines and obtuse tooth, movable spur slightly arcuate, basal tarsomere slender at base, carinate, with acute tubercle with two setae and a small tubercle at middle of tarsomere. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg somewhat widened.

Male genitalia ( Fig. 94 View FIGURES 85–96 ). Median lobe moderately arcuate at basal part, straight to apex, in lateral view widened at middle, extended acuminated to apex, base with basal oroficium in lateral view moderately distorted. Parameres robust, with well-developed apophyses, slightly distorted, hyaline at apex, each with two apical setae.

Female gonocoxites and epipleurite ( Fig. 120 View FIGURES 110–128 ). Basal gonocoxite with two long and two short setae, apical gonocoxite elongated, oval in cross section, with one long seta at basal third, with two very long setae apically. Epipleurite shaped like oval lappet, with one long seta.

Variation. In two of the paratypes, the supraorbital carinae are somewhat confluent with the frons, and the lateral margin of the pronotum is more or less trapezoid-shaped.

Distribution. Known from the type locality Tambarana on the island of Sulawesi.

Diagnosis. A small to medium sized fuscous species with trapezoid to cordate pronotum and intervals one to five at middle of the disk smooth and glossy. The similar species with smooth and glossy inner intervals of the elytron and a similar body size is C. circumreticulata sp. nov. from Borneo . It differs by the excised labrum which is in addition seven-setose. Moreover, the pronotum is distinctly square to cordate-shaped and the elytra are more slender and subparallel.

Etymology. The name refers to the elytra with its opaque appearance due to the reticulation on the lateral intervals, basally, and apically (Latin: circum = surrounded by; opacus, a, um = dull, darken, like with shadow).