Clivina caputopaca, Balkenohl, 2025

Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), Zootaxa 5592 (1), pp. 1-70 : 39-41

publication ID	https://doi.org/10.11646/zootaxa.5592.1.1
publication LSID	lsid:zoobank.org:pub:0E478171-152D-4A6F-9CB5-D9C9B5ABB744
DOI	https://doi.org/10.5281/zenodo.14953738
persistent identifier	https://treatment.plazi.org/id/03F487F9-7333-FF87-53D4-A577FB6DFE4E
treatment provided by	Plazi
scientific name	Clivina caputopaca
status	sp. nov.

Treatment

Clivina caputopaca sp. nov.

( Figs 24 View FIGURES 15–26 , 59 View FIGURES 53–67 , 92 View FIGURES 85–96 , 118 View FIGURES 110–128 )

Type material. Holotype: ♂, with labels and data: white, black printed “ MALAYSIA: Sabah Lahad Datu, Ulu Segama For. Res. Danum Valley Forest Centre 01°57’N 117°48’E, 450m alt xi.2005, 1° Forest coll. Mann, Slade & Villanueva” / “ Lowland mixed Dipterocarp frst around buildings at artificial light OUMNH-2005-062” / “ Clivina group helferi PUTZ. det. Balkenohl, 2020” / “desiderata ex Oxford” ( OUMNH). GoogleMaps

Paratypes: 1 ♀, same data as holotype ( CMBB) GoogleMaps ; 1 ♂, 3 ♀♀, with labels and data: white, black printed “ BORNEO , Sabah Danum Valley, Rainforest Lodge , 19.-10.X.2009, leg. U.+ H.J. BREMER” ( SMNS) .

Description. Measurements. Body length 3.84–4.53 mm (ẍ = 4.24 mm *), width 1.11–1.28 mm (ẍ = 1.22 mm *), ratio L/W of pronotum 0.86–0.88 (ẍ = 0.87), ratio L/W of elytra 1.91–1.93 (ẍ = 1.92*); (*n = 6). Colour glossy, fuscous to piceous. Supraantennal plate, lateral margin of pronotum and elytra slightly translucent.Antennae, mouthparts, intermediate and hind legs yellowish.

Head a seventh narrower than pronotum. Clypeus with central part straight, with indistinctly projecting rounded teeth laterally, separated from clypeal-wing by indistinct obtuse notch; wing slightly wider than teeth of clypeus, flatly rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse-angled slight notch, supraantennal plate with margin convex, clypeus indistinctly reflexed margined. Supraantennal plate extended posteriorly into an obtuse keel, separated from wing by indistinct furrow. Clypeus and frons forming continuous slight convexity, with confluent densely scattered small punctures and isodiametric reticulation, flat clypeal-furrow nearly not traceable. Frons with flat pore at middle, separated from supraantennal plate and supraorbital carina by flat reticulated furrow, with clypeal seta in front; supraorbital carinae moderately developed, with two supraorbital setigerous punctures at middle of eye and posterior gena-level. Neck constriction nearly closed, formed by a slight step from frons and reticulation posteriorly. Eye prominent, gena small but distinct. Antenna elongate, reaching over base of elytron. Labrum slightly excised anteriorly, finely isodiametrically reticulated, seven-setose. Mentum oval acuminated, with isodiametric reticulation, median tooth distinctly lower than lobe, acute at apex.

Pronotum ( Fig. 59 View FIGURES 53–67 ) with disk extraordinary flattened in lateral view, in frontal view laterally moderately convex, depressed at median line. Shape sub-cordate, wider than long. Reflexed lateral margin nearly smooth, with fine crenulation visible at 100-fold, convex at middle, widest at middle; anterior and posterior angles distinct, posterior one with distinct projecting tooth; lateral channel narrow, continuing up to base as sharp, straight margin. Median line complete, depressed, engraved at middle, joining anterior transverse line. Anterior transverse line distinct, complete. Disk covered with micro and medium sized punctures, with rough transverse wrinkles laterally, with isodiametric reticulation at base, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 45°. Basal channel on flange somewhat widened, rugose, with sharp carina.

Elytron ( Fig. 24 View FIGURES 15–26 ) with disk distinctly flattened in anterior half in lateral view, moderately convex in frontal view. Shape of elytra elongate, about two times longer than wide, with lateral margin nearly straight at middle, indistinctly diverging, maximum width posterior middle. Humerus rounded off. Base distinctly retracted. Reflexed lateral margin indistinctly crenulate behind humerus (120-fold). Lateral channel slightly narrower at middle. Row of setigerous punctures in the lateral channel nearly regular in size and distance, scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, impunctate, one to four free at base, five and six joining at humerus, one to four ending free at apex, five and six joining weakly apically. Intervals slightly to moderately convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven completely flat at apex, eight step-like carinate at apex. Surface of interval one to five at middle of disk smooth and glossy, all other intervals with distinct isodiametric reticulation, apically third of elytron reticulated as well as base.

Hind wing fully developed.

Lower surface. Proepisternum isodiametrically reticulated, with fine wrinkles. Sternite of abdomen with isodiametric reticulation. Last visible sternite with the two marginal setigerous punctures widely separated, with moderately narrow projection posteriorly at middle where surface is smooth, with narrow marginal furrow at middle (in both sexes).

Legs. Protibia moderately slender, with indistinct longitudinal reticulation, with sulcus dorsally, with three slender acute spines and small but distinct tooth, movable spur moderately arcuate, basal tarsomere slender at base, carinate, with acute tubercle with two setae. Intermediate tibia with nine tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg somewhat widened.

Male genitalia ( Fig. 92 View FIGURES 85–96 ). Median lobe distinctly arcuate at basal part, slightly arcuate up to apex, in lateral view widened at middle, short acuminated to apex. Basal oroficium not distorted. Parameres slender, with small apophyses, slightly distorted, hyaline at apex, each with one minute setae apically.

Female gonocoxites and epipleurite ( Fig. 118 View FIGURES 110–128 ). Basal gonocoxite with three long setae, apical gonocoxite elongated, oval in cross section, flattened in basal part, with one long seta at base, one long seta at middle, with two very long setae apically. Epipleurite relatively wide, asetose.

Variation. In two of the paratypes, the reticulation on the head is not as distinct developed on the top of the supraantennal plates and in the posterior part of the frons. Among the six specimens, the lateral margin of the pronotum varies in its convexity, and the micro and medium sized punctures on the disk vary in number.

Distribution. Known from Sabah on the island of Borneo .

Diagnosis. A small sized fuscous to piceous species with sub-cordate pronotum, intervals one to five at middle of the disk smooth and glossy, and with seven-setose labrum. The most similar species are C. circumopaca sp. nov. from Sulawesi and C. circumreticulata sp. nov. from Borneo , both having as well the middle of the elytra smooth and glossy. C. caputopaca sp. nov. differs from both of them mostly by the pronotum with its depressed median line, the elytron with interval seven completely flat and interval eight developed as step-like carina, the slightly developed clypeal-wings and clypeal-notches, the reticulated head, and the smaller size.

Etymology. The name refers to the head (Latin: caput = head) with its intense cover of reticulation giving the surface a dull opaque appearance (Latin: opacus, a, um = darken, dull, like with shadow).