Clivina punctaticollis, Balkenohl, 2025

Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), Zootaxa 5592 (1), pp. 1-70 : 50-52

publication ID	https://doi.org/10.11646/zootaxa.5592.1.1
publication LSID	lsid:zoobank.org:pub:0E478171-152D-4A6F-9CB5-D9C9B5ABB744
persistent identifier	https://treatment.plazi.org/id/03F487F9-7324-FF9C-53D4-A402FD3AFD36
treatment provided by	Plazi
scientific name	Clivina punctaticollis
status	sp. nov.

Treatment

Clivina punctaticollis sp. nov.

( Figs 33 View FIGURES 27–37 , 68 View FIGURES 68–72 , 100 View FIGURES 97–104 , 126 View FIGURES 110–128 )

Type material. Holotype: ♂, with labels and data: white, black printed “ SARAWAK ( Borneo ), ca 40 km SE KAPIT 03.1994, J. Kodada leg.” ( NHMW) . Paratypes: 1 ♂, same data as holotype ( CMBB) ; 1 ♀, with labels and data:

white, black printed “ BRUNEI: Temburong Kuala Belalong FSC, 60-300m ” / “ Clivina helferi group P.Bulirsch det. 2013” ( NHMW). 1 ♀ , with labels and data: white, black printed “ BORNEO BRUNEI Temburong Kuala Belalong R.Borcherding leg.” / “VI / VII 1995 KBFSC” / “ Clivina nr. wallacei PUTZ. det. Balkenohl IX.96” / “COLL. WRASE BERLIN” / “COLL. WRASE GUSOW-PLATKOW GERM.” ( CDWGP) .

Remark. In the female paratype ( CDWGP), the right elytron is missing.

Description. Measurements. Body length 4.03–4.54 mm (ẍ = 4.28 mm *), width 1.21–1.34 mm (ẍ = 1.27 mm *), ratio L/W of pronotum 0.83–0.87 (ẍ = 0.85*), ratio L/W of elytra 1.85–1.89 (ẍ = 1.87*); (*n = 4). Colour glossy, piceous. Supraantennal plate and clypeal-wing translucent. Antennae, mouthparts, intermediate and hind legs yellowish.

Head a sixth narrower than pronotum. Clypeus with central part straight, with slightly projecting teeth laterally, separated from clypeal-wing by obtuse notch; wing well visible, rounded, not as projecting as clypeal-teeth, separated from supraantennal plate by obtuse notch, supraantennal plate with margin laterally obtuse-angular, clypeus, wing, and supraantennal plate indistinctly reflexed margined. Supraantennal plate extended posteriorly into a convex keel, separated from wing by furrow. Clypeus slightly convex, with scattered small punctures, clypeal-furrow indistinct, flattened. Frons relatively wide, slightly convex, with pore at middle, with scattered small punctures, separated from supraantennal plate and supraorbital carina by wide furrow, with clypeal seta in front; supraorbital carinae moderately long, narrow, with two supraorbital setigerous punctures at middle of eye and distinctly posterior end of the gena-level. Neck constriction closed, formed by a step from frons. Eye well developed but somewhat flattened, gena small. Antenna elongate, just reaching up to base of elytron. Labrum excised anteriorly, isodiametrically reticulated, seven-setose. Mentum isodiametrically reticulated, lobe slightly hollowed out, laterally reflexed margined, bilaterally tapered to acute apex, median tooth acute, distinctly lower than lobes, clefts small.

Pronotum ( Fig. 68 View FIGURES 68–72 ) with disk distinctly flattened in lateral view, slightly convex in frontal view. Shape square to cordate, wider than long. Reflexed lateral margin smooth, nearly parallel, indistinctly convex, widest at middle; anterior angle distinctly marked, posterior angle distinctly marked by tooth; lateral channel moderately narrow, isodiametrically reticulated, continuing up to base as narrow, indistinctly convex margin. Median line engraved, complete, joining anterior transverse line. Anterior transverse line distinct, complete. Disk nearly completely covered with medium sized partly connected punctures, with some transverse wrinkles laterally, with isodiametric reticulation laterally and at base, with posterior setigerous puncture removed from margin by diameter of pore. Lateral margin from posterior angle to flange of base slanted with angle of around 40°. Basal channel on flange regularly developed, rugose, with sharp carina.

Elytron ( Fig. 33 View FIGURES 27–37 ) with disk distinctly flattened in lateral view, slightly to moderately convex in frontal view. Shape of elytra elongate, about two-fifth longer than wide, with lateral margin nearly straight at middle but slightly diverging, maximum width posterior middle. Humerus rounded but distinct. Reflexed lateral margin slightly crenulate behind humerus. Row of setigerous punctures in the lateral channel regular in size, slightly more distant at middle. Scutellar striole distinct; setigerous tubercle at base of first stria distinct, with distinct tubercle at base of third interval, basal declivity perpendicularly falling to pedunculus, with isodiametric reticulation. Striae moderately deep, nearly impunctate, one to four free at base, five and six joining at humerus, one to six ending free at apex. Intervals slightly convex, interval three with four setigerous punctures joining third stria, interval five joining lateral margin at humerus as short carina, interval seven flattened at apex, eight carinate at apex. Surface of interval one to six glossy, interval eight and half of width of interval seven with distinct isodiametric reticulation, apical fifth of elytron isodiametrically reticulated on all intervals.

Hind wing fully developed.

Lower surface. Proepisternum isodiametrically reticulated, without wrinkles. Sternite of abdomen with isodiametric reticulation, scalier medially. Last visible sternite with the two marginal setigerous punctures widely separated, with slight projection posteriorly at middle where it is flattened, with narrow marginal furrow at middle (in both sexes).

Legs. Protibia slender, with longitudinal reticulation, with sulcus dorsally, with three spines, movable spur distinctly curved, basal tarsomere slender at base, carinate, with acute tubercle and two setae at middle of tarsomere. Intermediate tibia with eight tubercles at the lateral side, with slender spur near apex, all setae furnished. Tarsomeres of front leg widened.

Male genitalia ( Fig. 100 View FIGURES 97–104 ). Median lobe distinctly arcuate basal part and at middle, slightly arcuate in apical half, apically regularly acuminate, somewhat flattened. Parameres moderately slender, with small apophysis, slightly distorted, hyaline apically, asetose.

Female gonocoxites and epipleurite ( Fig. 126 View FIGURES 110–128 ). Basal gonocoxite with two long setae, apical gonocoxite elongated, oval in cross section, with one long seta at base and one long seta above middle, with two very long setae apically. Epipleurite triangle-shaped, with one long seta arising from tubercle, with two medium sized setae.

Variation. In one of the paratypes, the neck constriction is indistinctly closed at middle. In the paratypes, the striae three and four and five and six of the elytron are more or less indistinctly joining at apex, and the extend of the reticulation on interval seven varies.

Distribution. Known from Sarawak and Brunei on the island of Borneo .

Diagnosis. A small sized piceous species with cordate to square pronotum and intervals seven and eight of the elytron reticulated. The most similar species C. bilobata sp. nov. differs by the labrum which is distinctly bilobate and excised, the pronotum with the indistinct posterior angle and lateral teeth, its more convex lateral margin, and the nearly smooth disk. In addition, C. bilobata sp. nov. is a larger species. Certain similarities can also be observed in C. margineserrata sp. nov., but that species shows a straight labrum, a pronotum with lateral margin distinctly crenulated, the lateral channel wide, and a widened basal channel. In addition, all intervals are reticulated.

Etymology. The name refers to the pronotum (Latin: collum = pronotum) with its intense cover of medium sized punctures (Latin: punctatus = with punctures).