Clivina neglecta, Balkenohl, 2025

Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), Zootaxa 5592 (1), pp. 1-70 : 56-63

publication ID	https://doi.org/10.11646/zootaxa.5592.1.1
publication LSID	lsid:zoobank.org:pub:0E478171-152D-4A6F-9CB5-D9C9B5ABB744
persistent identifier	https://treatment.plazi.org/id/03F487F9-7322-FF91-53D4-A739FE48F86D
treatment provided by	Plazi
scientific name	Clivina neglecta
status	sp. nov.

Treatment

Clivina neglecta sp. nov.

( Figs 37 View FIGURES 27–37 , 72 View FIGURES 68–72 , 104 View FIGURES 97–104 )

Type material. Holotype: ♂, with labels and data: white, black printed “ Sumatra Mjöberg” / “Kota Tjane” / “ F. van Emden Bequest. B.M. 1960-129.” / handwritten in black ink and black printed “Gesch. 8. 1938 von Andrewes” / “ Clivina integra Andr. H.E.Andrewes det. ( NHMUK).

Remark. The specimen is slightly abraded. Five terminal antennomeres at the left side and some frontal tarsomeres are missing, and the right front femur is broken. At the left hind leg, the three terminal tarsomeres are missing.

Description. Measurements. Body length 4.44 mm, width 1.37 mm, ratio L/W of pronotum 0.91, ratio L/W of elytra 1.82. Colour glossy, ferruginous. Supraantennal plate slightly brightened. Antennae, mouthparts and legs yellowish.

Head a quarter narrower than pronotum. Clypeus straight anteriorly, with its lateral teeth rounded and indistinctly projecting; wing not prominent, rounded, slightly less protruding than clypeus, separated from clypeus by small notch; supraantennal plate convex laterally, straight in anterior half; clypeus reflexed margined. Supraantennal plate extended up to mid-eye level, ending at mid-eye level as blunt knob, separated from wing by indistinct flat notch, separated from supraorbital carina by deep moderately wide furrow. Clypeus with transverse blunt horseshoe-like elevation, separated from frons by flat but distinct furrow. Frons moderately convex, smooth. Supraantennal plate separated from clypeus and frons by moderately wide furrow continuing posteriorly up to mid-eye level as moderately wide supraorbital furrow. With blunt distinct supraorbital carina at posterior half of eye. With two supraorbital setigerous punctures situated at mid-eye level and posterior gena-level. Neck constriction developed as step, distinct laterally, indistinct and flat at middle, laterally punctured posteriorly. Eye convex but somewhat flattened. Gena distinct, enclosing eye posteriorly as small stripe. Antenna somewhat shortened in comparison to body size, just reaching over posterior setigerous punctures of pronotum, antennomeres four to ten elongate (L/W around 1.88). Labrum slightly excised, isodiametrically reticulated, seven-setose. Mandible of moderate size. Mentum at middle with isodiametric reticulation, lateral lobe with fine coreaceate-like structure, with oblique carinae, lateral margin nearly straight anteriorly, obtuse angulated anteriorly, convex medially, median tooth widened apically, slightly higher as lateral lobes.

Pronotum ( Fig. 72 View FIGURES 68–72 ) with disk slightly convex in lateral view, in frontal view moderately convex at middle and distinctly laterally. Subsquare, wider than long, resembling to sub-globosity in dorsal view. Anterior margin excised. Reflexed lateral margin nearly smooth, with indistinct microscopic notches, distinctly convex, widest at middle; anterior and posterior angles distinctly rounded-off; tooth at posterior angle nearly integrated in rounding of posterior angle. Lateral channel relatively narrow, reticulated, margin from posterior angle to base slanted by angle of about 45°, indistinctly convex, flange short but distinct, basal channel and carina of flange of regular size. Median line distinct, crenulated, joining anterior transverse line, joining base; anterior transverse line complete. Surface with short impression bilaterally, with few punctures in basal half, basal declivity isodiametrically reticulated. Elytron ( Fig. 37 View FIGURES 27–37 ) with disk flattened in anterior half in lateral view, regularly convex in frontal view. Outline long oval, distinctly less than twice as long as wide, with maximum width behind middle. Humerus distinctly marked, formed by lateral margin which joins stria five laterally by a very short carina. Reflexed lateral margin nearly smooth posterior humerus. Lateral channel moderately wide. Scutellar striole of moderate length, not deep; with distinct setigerous puncture at base of first stria, with minute tubercle at base of third interval. Striae moderately deep, indistinctly punctate, one to four free at base, five and six joining at humerus, striae one and two running up to apex, three and four, and five and six joining apically. Intervals moderately convex, eights flattened, seven and eight carinate apically. Third interval with four setigerous punctures adjoining third stria. Surface of intervals smooth and glossy on disk, with distinct isodiametric reticulation at basal declivity, at apical fifth, and on interval eight.

Hind wing fully developed.

Lower surface. Proepisternum with distinct isodiametric reticulation, with transverse wrinkles laterally in posterior part. Sternites of abdomen with isodiametric reticulation laterally, shingle-like and transverse at middle. Abdominal sternum VII with the two anal setigerous punctures widely separated, at middle slightly projecting posteriorly where it is flattened, with narrow marginal furrow at middle.

Legs. Protibia with three spines of moderate length and a small knob basally, slightly sulcate dorsally, movable spur wide, slightly arcuate, surface with longitudinal reticulation. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with seven setae furnished tubercles above protuberance, with few fine setae at the inner side. Front leg with tarsomeres of about same width as other tarsomeres, first tarsomere elongated, with carina baso-laterally, laterally with acute tubercle furnished with two short setae.

Male genitalia ( Fig. 104 View FIGURES 97–104 ). Median lobe arcuate in basal third, straight at middle, conspicuously arcuate in apical fifth including lamella, base with basal oroficium in lateral view at axis of median lobe. Lamella in lateral view duck-bill shaped. Parameres slender apically, spatulate, and hyaline in apical third, asetose (at 500-fold).

Female gonocoxites and epipleurite unknown.

Variation unknown.

Distribution. Known from the type locality Kota Tjane in the North of Sumatra.

Diagnosis. A small sized ferruginous species with interval eight of the elytron flattened and reticulated. It differs mainly from the most similar species C. integra Andrewes and C. similata sp. nov. by its subsquare shape of the pronotum with all angles distinctly rounded, the flattened eyes, the indistinct flattened neck constriction, and the median lobe of the male aedeagus in dorsal view which is conspicuously arcuate in apical fifth including the lamella, and the duck-bill shaped lamella in lateral view.

Etymology. The name is derived from the Latin expression ‘neglectus est’ which means ‘the overlooked one’, because the specimen of this species was sorted under type and other material of C. integra Andrewes.

Specimen incertae sedis

The following specimen could not be interpreted by its unclear character states, and severe damages: 1 spec., Sumatra / Coll. Kraatz / Andrewes det. / Clivina integra Andr. redet. K. Kult 44 / COLLECTIO KAREL KULT COLL.A.DOSTAL, 1999 (CADW). The specimen is glued at several parts, the right front leg is missing, and it was received with an empty hind body. Therefore, information on the genitalia is completely missing. It is not conspecific with the type and other material of C. integra . The data of this specimen have not been included under the determined material.

Concluding remarks:

On the basis of the limited material investigated, taxa of the Clivina westwoodi -species group seem to be mainly distributed in the eastern part of the Oriental region including New Guinea / Papua. It looks like that in the many islands a rich Clivinini fauna has been evolved with a high degree of endemism on some of the islands like for example in Borneo , Sulawesi, and New Guinea / Papua. Such high degree of endemism in Carabidae was also reported by Liebherr (2017) for the genus Mecyclothorax Sharp in New Guinea / Papua. This hypothesis is supported by the ascertainment that the species are not reported for Australia in the recently published profound revisions of the genus Clivina and allied genera by Baehr (2008, 2015, 2017) and are obviously not occurring there according to current knowledge.

At the first glance, the number of 637 specimens totally investigated is seemingly high. In reality, it is a small cut-out only. Many of the islands are only explored by random sampling, if at all, e.g. the Philippine Island with its about 7000 islands, Sulawesi, or New Guinea / Papua. Often, the samplings exist of few specimens, sometimes only one specimen of a species was found .

Moreover, the morphological differences are in some cases not large respectively the characters and combination of characters were not known in this specificity from other regions. So, differential diagnoses based on the habitus only reach limits. Fortunately, characters of the male and female genitalia are distinct in all the cases evaluated. A good example is the pair C. westwoodi and C. wallacei wallacei which are similar but show a different external female reproductive tract. Taking the explanations regarding the collection locality and the assumption of Darlington (1962) into account, the hypothesis that C. wallacei wallacei occurs in Sulawesi and not in New Guinea / Papua might be probably true. The new finds on Sulawesi support this. The lectotype of C. westwoodi from New Guinea is the only specimen actually available and needs such confirmation as well.

Therefore, the revision is seen as an initial stage, and exact conclusions on the distribution cannot be drawn at the current point in time.