Clivina convexaquadrata, Balkenohl, 2025

Balkenohl, Michael, 2025, On Asian Clivinini. Revision of the westwoodi-species group of the genus Clivina Latreille, 1802 (Coleoptera, Carabidae, Clivinini), Zootaxa 5592 (1), pp. 1-70 : 14-15

publication ID	https://doi.org/10.11646/zootaxa.5592.1.1
publication LSID	lsid:zoobank.org:pub:0E478171-152D-4A6F-9CB5-D9C9B5ABB744
DOI	https://doi.org/10.5281/zenodo.14953688
persistent identifier	https://treatment.plazi.org/id/03F487F9-7318-FFA1-53D4-A20BFAECFA5E
treatment provided by	Plazi
scientific name	Clivina convexaquadrata
status	sp. nov.

Treatment

Clivina convexaquadrata sp. nov.

( Figs 5 View FIGURES 3–14 , 40 View FIGURES 38–52 , 75 View FIGURES 73–84 , 107 View FIGURES 105–109 )

Type material. Holotype: ♂, with labels and data: white, black printed “ BORNEO , Sabah, Danum Valley, Raiforest Lodge , 19.-20.X.2009. leg. U. + H.J. BREMER” ( SMNS).

Paratypes: 1 ♀, 1 spec., same data as holotype ( CMBB) ; 1 spec., “ Borneo – MALAYSIA: Sabah: Mt.Kinabalu Park vic. Serinsim sub-station” / “ 6°12’N 116°33’E L.F. 180-250m 30.VIII.1998 D.Bartsch & C.Häuser leg.“ / „ Clivina helferi group det Balkenohl 2000” ( SMNS) GoogleMaps ; 1 ♂, “ BORNEO , SABAH, NW., Kota Marundu, Taritipan env., 13.v.1999, M.Sniźek leg.” / “helferi Gr. ” ( CADW) ; 1 spec., “ Malaysia, Sabah, Batu Punggul Resort env. 24.VI.-1.VII.1996, 11f, intercept trap ” ( NHMW) ; 1 ♂, “ Sarawak, Kapit dist. Rumah Ugap vill., Sut riv., 3.-9. 3. 1994 J. Horák leg.” ( CADW) ; 1 spec., “ BORNEO , Brunei 2010 Temburong Prov. 14.-15.ii. Kuala Belalong F. stn., at light, BMNH (E) 2010-15” ( NHMUK) ; 1 ♀, 3 specs, “ SARAWAK ( Borneo ), ca 25 km E KAPIT, III. 1994, Kodada leg.” ( NHMW) ; 1 spec., same data but “ca 40 km SE KAPIT” ( NHMW) ; 1 spec., “ MALAYSIA – Sabah prov. Banjaran Crocker Mts. GUNUNG ALAB peak 30.IV.-27.V.1996, 1650-1800m M. Štrba & R. Hergovitis leg.” / “VIII. helferi Gr. det. Dr. A. Dostal 2000”. This specimen was received with an empty abdomen. ( CADW) ; 1 ♂, “ BORNEO : SABAH Bingkor N Keningau 400-500 m, 19.-20.XI.1996 leg. W. SCHAWALLER” / “ Clivina group helferi, nr. wallacei PUTZ. det. Balkenohl, 1998” ( SMNS) .

Description. Measurements. Body length 6.31–7.19 mm (ẍ = 6.59 mm *), width 1.91–2.28 mm (ẍ = 2.05 mm *), ratio L/W of pronotum 0.82–0.85 (ẍ = 0.84*), ratio L/W of elytra 1.77–1.88 (ẍ = 1.81*); (*n = 10). Colour glossy, fuscous. Supraantennal plate with margin and clypeal-wing translucent. Antennae, mouthparts, intermediate and hind legs yellowish. Elytron with tip of apex brightened.

Head a quarter narrower than pronotum. Clypeus straight anteriorly, with its lateral teeth slightly projecting; wing of moderate size, not more protruding anteriorly than clypeus, distinctly separated from clypeus by notch; supraantennal plate convex laterally, with straight part in anterior half; clypeus, wing, and supraantennal plate finely reflexed margined. Supraantennal plate extended up to mid-eye level, separated from wing by distinct notch, separated from supraorbital carina by a step. Clypeus transverse, with transverse flat V-like elevation, separated from frons by flat furrow. Frons moderately convex, with central pore at middle, with scattered small punctures. Supraantennal plate separated from clypeus and frons by moderately wide furrow continuing posteriorly up to mid-eye level as moderately wide supraorbital furrow. With blunt distinct supraorbital carina at posterior half of eye. Posterior keel of supraorbital plate and supraorbital carina separated by distinct step. With two supraorbital setigerous punctures situated at mid-eye level and at the posterior end of eye. Neck constriction developed by punctures and distinct step, step pointed posteriorly at middle. Eye distinctly hemispherical projecting laterally. Gena small. Antenna moderately long, not reaching base of pronotum, antennomeres four to ten elongate (L/W around 1.55). Labrum nearly straight, isodiametrically reticulated, seven-setose. Mandible of moderate size. Mentum with lateral lobe rectangular anteriorly, flat, isodiametrically reticulated, median tooth as high as lateral lobes, hollowed out, slightly rounded apically.

Pronotum ( Fig. 40 View FIGURES 38–52 ) with disk somewhat flattened but slightly convex in lateral view, in frontal view flattened at middle and moderately convex laterally. Quadrate, wider than long. Anterior margin slightly excised. Reflexed lateral margin smooth, slightly convex at middle, attenuating in anterior half, widest in posterior third; anterior angle distinct but rounded; posterior angle marked by laterally distinctly projecting tooth. Lateral channel wide (lateral view), reticulated, basal part of lateral margin from posterior angle to flange of base slanted with angle of slightly more than 20°, indistinctly convex, flange short but distinct, basal channel moderately developed, basal carina sharp. Median line moderately narrow, joining anterior transverse line and base; anterior transverse line narrow, complete. Surface with microscopic punctures, bilaterally with three group of punctures in basal half, arrangement of puncture-groups resembles to an indistinct widened letter Y, base isodiametrically reticulated.

Elytron ( Fig. 5 View FIGURES 3–14 ) with disk flattened in anterior half in lateral view, regularly convex in frontal view. Outline long oval, distinctly less than twice as long as wide, with maximum width behind middle. Humerus rounded, formed by lateral margin which joins stria five. Reflexed lateral margin subcrenulated posterior humerus. Lateral channel moderately wide. Scutellar striole of moderate length, not deep; with distinct setigerous puncture at base of first stria, with small tubercle at base of third interval. Striae moderately deep, with indistinct punctures, one to four free at base, five and six joining at humerus, seven shortened behind humerus, striae one and two running up to apex, three and four, and five and six just not joining apically. Intervals moderately convex, eight carinate apically. Third interval with four setigerous punctures adjoining third stria. Surface of intervals smooth and glossy on disk, with distinct isodiametric reticulation at base, at apical tip, and on interval eight, with indistinct reticulation at interval seven.

Hind wing fully developed.

Lower surface with proepisternum completely covered with distinct isodiametric reticulation, with some transverse wrinkles. Sternites of abdomen isodiametrically reticulated, transverse at middle, the three middle segments with flat impression laterally. Abdominal sternum VII with the two anal setigerous punctures widely separated, at middle slightly projecting posteriorly where it is flattened, with narrow marginal furrow at middle (in both sexes).

Legs. Protibia with three spines of moderate length and a small but distinct one basally, sulcate dorsally, movable spur nearly straight, surface with nearly isodiametric reticulation. Mesotibia with distinct and moderately long protuberance preapically with acuminate apex and with its seta inserted laterally, with nine setae furnished tubercles above protuberance, with few fine setae at the inner side. Front leg with tarsomeres slightly widened, first tarsomere elongated, slender and with carina basally, laterally with three setae furnished tubercles.

Male genitalia ( Fig. 75 View FIGURES 73–84 ). Median lobe with nearly straight part at middle, curved basally and slightly at root of apical lamella, apex with flattened duck bill-like elongated lamella. Parameres moderately wide, acuminated and slightly distorted in apical third, apex asetose. Female gonocoxites and epipleurite ( Fig. 107 View FIGURES 105–109 ). Gonocoxites scapiform, extraordinary slender, less sclerotized, not fused, completely movable; basal gonocoxite 1 elongated, with three setae, apical gonocoxite 2 slender, distinctly elongated, indistinctly curved towards apex, cylindrical in apical half, with two long setae at apex, with two moderately long setae at lateral margin at middle. Epipleurite with two setae distant from each other.

Variation. The reticulation on interval seven is more or less expressed. Specimens from Sarawak exhibit a strip of reticulation, only. The flat clypeal-furrow on the head is slightly rugose in some specimens, and the clypeus is more or less elevated. The specimen from Bingkor is rather small.

Distribution. Known from the island of Borneo : Sabah, Brunei, Sarawak.

Diagnosis. A larger sized dark fuscous species with square pronotum and interval seven and eight of the elytron reticulated. It differs mainly from the most similar species C. quadrataconvexa Balkenohl by the anteriorly distinctly attenuated pronotum with laterally projecting sharp teeth of the posterior angle, the wider basal channel, and with intervals seven and eight of the elytron reticulated. It differs from C. tuberculifer sp. nov. mainly by the elytron with small tubercle at the base of interval three, the more intense reticulation on intervals seven and eight, the smaller eyes, and the smaller size.

Etymology. The name refers to the square pronotum which is in frontal view laterally distinctly convex, and is expressed by the Latin adjective convexus (= arched) and quadratus (= square) in the feminine genus.