Borbo gemella (Mabille, 1884), 2023

Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina & Grishin, Nick V., 2023, Butterfly classification and species discovery using genomics, The Taxonomic Report of the International Lepidoptera Survey 11 (3), pp. 1-94 : 80

Afrogegenes Jong & Coutsis, 2017 and Torbenlarsenia Kemal & Koçak, 2020 are subgenera of Gegenes Hübner, 1819

Nuclear genome trees point to a strongly supported clade that is sister to Borbo Evans, 1949 (type species Hesperia borbonica Boisduval, 1833 ) and includes type species of three genera: Gegenes Hübner, 1819 Hesperia hottentota Latreille, 1824 ), and Torbenlarsenia Kemal & Koçak, 2020 (type species Hesperia holtzi Plötz, 1882 ) in addition to several species currently in Borbo Evans, 1949 (type species Hesperia borbonica Boisduval, 1833 ) ( Fig. 50). However, within this clade, strongly supported major subclades are absent, and Torbenlarsenia as currently circumscribed ( Fan et al. 2016) is not monophyletic. Moreover, Torbenlarsenia gemella (Mabille, 1884) (type locality in Madagascar) is not in this clade but belongs to Borbo , where we transfer it as Borbo gemella (Mabille, 1884) , stat. rev. This whole clade with Gegenes is at the genus level in the tree and splits into five confident subclades, three of which represent named genus-group taxa: we propose to treat Afrogegenes Jong & Coutsis, 2017 , stat. rev. and Torbenlarsenia Kemal & Koçak, 2020 , stat. rev. as subgenera of Gegenes Hübner, 1819 . The remaining two subclades do not have names and are described as new subgenera below. Finally, to restore the monophyly of Borbo , we transfer several species to the subgenus Torbenlarsenia , forming new combinations: Gegenes (Torbenlarsenia) cottrelli (Larsen, 2013) , comb. nov., Gegenes (Torbenlarsenia) fallax (Gaede, 1916) , comb. nov., Gegenes (Torbenlarsenia) fanta ( Evans, 1937) , comb. nov., Gegenes (Torbenlarsenia) micans ( Holland, 1896), comb. nov., and Gegenes (Torbenlarsenia) sirena ( Evans, 1937) , comb. nov.