Sape Mabille, 1891

Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina & Grishin, Nick V., 2023, Butterfly classification and species discovery using genomics, The Taxonomic Report of the International Lepidoptera Survey 11 (3), pp. 1-94 : 67-70

Sape Mabille, 1891 is a genus distinct from Sarangesa F. Moore, [1881]

Genomic tree reveals that Sape lucidella Mabille, 1891 , the type species of the genus Sape Mabille, 1891 currently treated as a junior subjective synonym of the genus Sarangesa F. Moore, [1881] (type species Sarangesa albicilia F. Moore, [1881] ) is not monophyletic with it (Fig. 44 magenta). We find that Eretis Mabille, 1891 (type species Eretis melania Mabille, 1891 ) renders Sarangesa paraphyletic. Therefore, to restore the monophyly, we propose that Sape Mabille, 1891 , stat. rest. is a genus distinct from Sarangesa F. Moore, [1881] .

Cupithina Grishin, new subtribe

http://zoobank.org/ 905E8110-34EC-4CB3-A0C0-3F9002414D3E

Type genus. Cupitha F. Moore, 1884 .

Definition. The tribe Astictopterini Swinhoe, 1912 splits into two clades containing approximately the same number of genera, each supported by 100% of partitions (see Fig. 1 in Zhang et al. (2023b) and Figs. 45, 46). Although neither clade is very prominent (branches separating them are not particularly long), both are most strongly supported. Therefore, to bring order to the genera in Astictopterini , we divide the tribe into two subtribes. The clade with the type species of the tribe contains generally larger species with more robust bodies and corresponds to the nominotypical subtribe. The second clade, with Cupitha (type species Cupitha tympanifera F. Moore, 1884 , which is a junior subjective synonym of Pamphila purreea Moore, 1877 ), consists of smaller and frequently brighter patterned species and corresponds to the subtribe without an available name. This new and mostly Afrotropical subtribe is distinguished from its relatives by a combination of the following characters ( Evans 1937; Evans 1949): palpi either porrect with convergent 3 rd segment or erect with thin and erect 3 rd segment; antennal club with pointed apiculus, either obtuse or hooked; caudal angle of hindwing discal cell not bent up: median vein and vein M 3 collinear, end of cell relatively straight, and vein 3A usually shorter than vein CuA 2; thorax not shaggy beneath; coxae and tibiae of hindlegs not densely fringed; valvae symmetrical, uncus typically narrowing to a point, ovate, terminal part narrow, frequently needle-like, uncus usually not expanded terminally and not hourglass-shaped. Most confidently identified by DNA and a combination of the following nuclear genomic base pairs is diagnostic: aly 1775.3.2:A50G, aly27.16.1:A638C, aly27.16.1:C644G, aly1603.20.3:G76C, aly1651.28.7:A167G.

Genera included. The type genus (i.e., Cupitha F. Moore, 1884 ), Acada Evans, 1937 , Acleros Mabille, 1885 , Andronymus Holland, 1896, Caenides Holland, 1896, Ceratricula Larsen, 2013 , Fresna Evans, 1937 , Gorgyra Holland, 1896, Gyrogra Lindsey & Miller, 1965 , Hollandus Larsen & Collins, 2015 , Hypoleucis Mabille, 1891 , Melphina Evans, 1937 , Melphinyet Larsen, 2012 , Noctulana Larsen, 2012 , Osmodes Holland, 1892, Osphantes Holland, 1896, Paracleros Berger, 1978 (see below), Paronymus Aurivillius, 1925 , Parosmodes Holland, 1896, Platylesches Holland, 1896, Rhabdomantis Holland, 1896, Semalea Holland, 1896, Teniorhinus Holland, 1892, Xanthodisca Aurivillius, 1925 (see below),

Zhang et al. (2023b) belong to the nominotypical subtribe Astictopterina .

Parent Taxon. Tribe Astictopterini Swinhoe, 1912 .