Grishin, Zhang & Cong & Shen & Song & Grishin, 2023

Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina & Grishin, Nick V., 2023, Butterfly classification and species discovery using genomics, The Taxonomic Report of the International Lepidoptera Survey 11 (3), pp. 1-94 : 70

Isocleros Grishin , new subgenus

http://zoobank.org/ 1109E9E2-43A8-422D-BF56-967608087687

Type species. Pamphila (?) mackenii Trimen, 1868 .

Definition. As discussed in the previous section, Acleros Mabille, 1885 (type species Cyclopides leucopyga Mabille, 1877 ) that includes Paracleros Berger, 1978 (type species Acleros biguttulus Mabille, 1889 ) splits into three prominent clades: the nominotypical Acleros , Paracleros , and the third clade not associated with any available genus-group names ( Figs. 45, 46). The third clade represents a new subgenus that keys to 41.B.(a) (excluding a 2) or 41.B.(b)(b 1) in Evans (1937) and is distinguished from its relatives by darker distal half of ventral hindwing or if hindwing is more uniform in color then dorsal forewing is with broadly white inner margin; tegumen with uncus are narrow in lateral view, uncus in dorsal view is either bulb-shaped or continuously narrows to a point. A combination of the following nuclear genomic base pairs is diagnostic: aly 2627.4.1:T63A, aly 2627.4.1:C69T, aly116.29.1:T2049A, aly116.29.1:G2814A, aly9673.4.1:A480G.

Etymology. The name of the other subgenus was formed by adding the prefix par - (i.e., besides, alongside) to the genus name. Similarly, we fuse the prefix iso - (i.e., similar, equal, the same) to the genus name. The name is a masculine noun in the nominative singular.

Species included. The type species (i.e., Pamphila (?) mackenii Trimen, 1868 ), Acleros bibundica Strand, 1913 , Acleros instabilis Mabille, 1889 , Apaustus olaus Plötz, 1884 , Acleros ploetzi Mabille, 1889 , and a new species described below.

Parent taxon. Genus Acleros Mabille, 1885 .