Lycaenidae

Zhang, Jing, Cong, Qian, Shen, Jinhui, Song, Leina & Grishin, Nick V., 2023, Butterfly classification and species discovery using genomics, The Taxonomic Report of the International Lepidoptera Survey 11 (3), pp. 1-94 : 30-31

Family Lycaenidae [Leach], [1815]

Liphyrinae Doherty, 1889 is a subfamily

Currently treated as a tribe within subfamily Miletinae Reuter, 1896 , Liphyrini Doherty, 1889 show corresponds to subfamilies ( Fig. 20). Therefore, we return this morphologically unique group to the status of a subfamily as originally proposed: Liphyrinae Doherty, 1889 , stat. rest.

Megalopalpina Grishin, new subtribe

http://zoobank.org/ 2A22AFFB-346D-4DCF-B256-C59DA7FCEFB1

Type genus. Megalopalpus Röber, 1886 .

Definition. Megalopalpus Röber, 1886 (type species Megalopalpus simplex Röber, 1886 ) forms a lineage sister to all other Miletini Reuter, 1896 and is more prominently separated from them than they are from each other ( Figs. 20, 21). Therefore, we propose that this lineage represents a subtribe. This new subtribe is diagnosed by a combination of the following characters, as given for the Megalopalpus section by Eliot (1973): uncus with tegumen plates triangular, with a unique for this group broad lobe-like triangular ventrally-pointed process, falces strongly curved, males lack secondary sexual characters, hindwing with precostal vein. A combination of the following nuclear genomic base pairs is diagnostic: cce64.1.4:G152C, cce5960.3.7:A41T, cce7884.1.3:A388A (not C), cce28693.2.4:G109G (not A), cce9880.2.2:A83A (not C), cce49.4.1:C595C (not T), cce16970.13.7:T187T (not G).

Genera included. Only the type genus.

Parent Taxon. Tribe Miletini Reuter, 1896 .

Tarakina Eliot, 1973 is a subtribe

Currently treated as a tribe in the subfamily Miletinae Reuter, 1896 , Tarakini Eliot, 1973 is sister to Spalgini Toxopeus, 1929 and shows less extensive genetic differentiation from them than typical for the tribal level ( Fig. 21). The Taraka Doherty, 1889 lineage has split from Spalgini at the same tree level as Megalopalpina subtrib. n. has split from Miletina Reuter, 1896 . Therefore, we place Tarakina Eliot, 1973 , stat. rev. as a subtribe in the tribe Spalgini , 1929.