Pteroscion peli (Bleeker, 1863)

Schwarzhans, Werner, 2023, Geology and stratigraphy of the Neogene section along the Oued Beth between Dar bel Hamri and El Kansera (Rharb Basin, northwestern Morocco) and its otolith-based fish fauna: a faunal inventory for the Early, Swiss Journal of Palaeontology (4) 142 (1), pp. 1-85 : 62

Pteroscion peli (Bleeker, 1863)

Figure 28i–l

1993 Pteroscion maroccanus —Schwarzhans: Fig. 289–292.

2003 Pteroscion peli (Bleeker, 1863) —Mendiola & Martínez: Fig. 9 :4–6, pl. 13, Fig. 289–291.

2003 Pteroscion guardamarensis —Mendiola & Martínez: Fig. 9 :7–9, pl. 12, Figs. 1–8 , pl. 15, Figs. 1–8 , pl. 16, Figs. 1–2 .

2003 Pteroscion sp. 1 —Mendiola & Martínez: pl. 12, Figs. 9–16 , pl. 16, Figs. 3 , 4 .

Material 46 specimens, Dar bel Hamri, Zanclean, figured specimens are photographs of holotype of P. maroccanus, SMF P 8599 ( Fig. 28h ) and paratypes, SMF P 8600

( Fig. 28i–k ).

Discussion I agree with Mendiola and Martínez (2003) and in the light of the many extant and fossil species they figured that P. maroccanus is synonymous with the extant P. peli . However, I also agree with the view of the reviewer of their manuscript, D. Nolf (as mentioned in their acknowledgements), that the species they described from the Pliocene of Spain as P. guardamarensis represent a further synonym of P. peli . Te species was found to be by far the most common sciaenid in the lower Pliocene rocks of Guardamar in southern Spain ( Mendiola & Martínez, 2003). Today, Pteroscion peli is distributed from Senegal to Namibia.

Figures

Fig. 28 Sciaenidae. a–d, Afroscion trewavasae Schwarzhans, 1993, Dar bel Hamri, Zanclean, b holotype, SMF P8226,a, c–d paratypes, SMF P 8227 (reversed).e Atractoscion cf. macrolepis Song, Kim & Kim,2017, SMF PO 101.296, Dar bel Hamri.f Umbrina canariensis Valenciennes, 1843, SMF PO 101.272 (reversed), Dar bel Hamri. g Pseudotolithus cf. typus Bleeker, 1863, SMF PO 101.270, Dar bel Hamri. h Pseudotolithus sp., SMF PO 101.271 (reversed),Dar bel Hamri. i–l Pteroscion peli (Bleeker, 1863),types of Pteroscion maroccanus Schwarzhans, 1993, Dar bel Hamri, Zanclean, h holotype, SMF P 8599, i–k paratypes, SMF P 8600 (k–l reversed)

Fig. 9 Pterothrissidae.a–e, Pterothrissus darbelhamriensis Schwarzhans,1981, Dar bel Hamri, Zanclean;a holotype, SMF P 6234;paratypes,b SMF 6235 (reversed), d SMF P 6237 (reversed), e SMF P 6236; c Asilah, coll.WWS. f–j Pterothrissus compactus Schwarzhans, 1981, Le Puget, southern France, Zanclean;f holotype, SMF P 6239;paratypes,g SMF P 6240, i SMF P 6243 (reversed); h, j (reversed), MGPT-PU 139453,Stazzano,Piedmont, Italy,Tortonian

Fig. 1 Regional map of northern Morocco outlining distribution of Neogene sediments. Schematized and simplified after the detailed map published by Saadi et al. (1980). The study areas highlighted are the Oued Beth outcrop section (red rectangle) and the Asilah outcrop (red asterisk)

Fig. 2 Geological map of the studied area from Dar bel Hamri to El Kansera resulting from a review of my master thesis (1977), and highlighting sections sampled.The location of the subsurface Sidi Fili fault and Nappe Prérifaine front are extrapolated from Tilloy (1952) and the postulated extension of the Kenitra-Sidi Slimane fault is from Zouhri et al. (2002)

Fig. 3 Photographs of selected section locations. a Base of section 2 at Sidi Mohamed ech Chleuh above river level. b, c Kef Nsour cliff (section 4); b panoramic view; c view of cliff top with Piacenzian sediments exposed. d, e Tortonian outcrop sequence near Sidi bou Sensal (section 7, and section 6 to the left in d); d panoramic view; e eroded basal Tortonian bioturbated mudstones at section 7 in the Oued Beth river bed; f panoramic view of upthrusted Jurassic rocks of the Jebel Kansera along the Oued Beth gorge

Fig. 4 Schematized outcrop-section along the Oued Beth right river bank based on studied sections

Fig. 5 Range chart of selected planktonic foraminifera observed in the study area.Ranges after mikrotax.org (2022) and Aze et al. (2011) and Maniscalco and Brunner (1998) for the Globorotalia margaritae plexus.Occurrences in studied sections and related regional biostratigraphic events mainly after Wernli (1988), Barhoun and Bachiri Taoufiq (2008) and Yousfi et al. (2013). Numbers refer to the sampled sections shown in Fig. 2

Fig. 6 Photoplate of selected planktonic foraminifera deemed of stratigraphic significance. Globorotalia margaritae primitiva: a Sidi Mohamed ech Chleuh (section 2), Zanclean level 2. Globorotalia margaritae margaritae: b, Kef Nsour (section 4), higher Zanclean; c, d, Sidi Mohamed ech Chleuh (section 2), Zanclean level 2;. Globorotalia margaritae evoluta: e, Kef Nsour (section 4), higher Zanclean. Globorotalia praemargaritae: f–h, Sidi bou Sensal (sections 6 and 7),Tortonian clay. Globorotalia cibaoensis?: i Sidi Mohamed ech Chleuh (section 2), Zanclean level 1. Globorotalia crassula: j Kef Nsour (section 4), basal Zanclean.Globorotalia juanai?: k Jebel Zebbouj (section 3), higher Zanclean.Globorotalia plesiotumida: l Sidi bou Sensal (section 7), basal Tortonian mudstone. Globorotalia saphoae: m Sidi bou Sensal (section 7), Tortonian clay.Globorotalia miotumida: n Sidi Mohamed ech Chleuh (section 2), Zanclean level 2 (presumed reworked). Globorotalia merotumida: o, p Dar bel Hamri (section 1), Zanclean, yellow sands of Dar bel Hamri (presumed reworked). Neogloboquadrina acostaensis: q Sidi bou Sensal (section 7),Tortonian clay; r Kef Nsour (section 4), basal Zanclean. Neogloboquadrina incompta: s Sidi bou Sensal (section 6), Tortonian clay.Neogloboquadrina dutertrei: t Jebel Zebbouj (section 3), higher Zanclean. Globigerinoides extremus: u Jebel Zebbouj (section 3), higher Zanclean. Sphaeroidinellopsis kochi: v Dar bel Hamri (section 1), coquina. Sphaeroidinellopsis seminulina: w, x Sidi bou Sensal (section 7),Tortonian clay. Sphaeroidinella dehiscens: y Jebel Zebbouj (section 3), higher Zanclean

Fig. 7 Photoplate of selected macrofossils from the Early Pliocene coquina of section 1, ca.1.5 km south of Dar bel Hamri. A rock sample is shown in n

Fig. 8 Conceptual structural cross-section through studied area based on Zizi (1996) and own field observations (scale 1:1). Subsurface thicknesses are extrapolated from sections of nearby wells published in Saadi et al. (1980) and Sani et al. (2006). Subsurface positions of not-exposed tectonic elements after Tilloy (1952), Zizi (1996), and Zouhri et al. (2002)