Flintoconis, Sziráki, 2007

Sziráki, Gy., 2007, Studies On Brucheiserinae (Neuroptera: Coniopterygidae), With Description Of The Second Genus Of The Subfamily, Acta Zoologica Academiae Scientiarum Hungaricae 53, pp. 231-254 : 238-239

Flintoconis gen. n.

Type species by present designation: Flintoconis gozmanyi sp. n.

Relatively large bodied and long winged ( Fig. 11 ) coniopterygids. Head rather small, and bent backwards tightly to the prothorax in usual position. (In this case it is hidden below the pronotum.) Postclypeus, upper part of labrum and the antennal sockets weakly chitinized, other parts of the head capsule well sclerotized, with a well developed, straight midfrontal-coronal apodeme ( Fig. 12 ). Structure of labial and maxillary palpi as it is usual in Coniopterygidae . Frons and vertex are not separated from each other. Antennae are more than fifty segmented, and about as long as the body. Pronotum large, wide, with well sclerotized edges, and with a median and two transverse apodemes ( Fig. 13 ). The anterior and posterior edges of pronotum are tucked up.

Wings are very long and rather narrow; the fore wings are about (or almost) two times as long as the body. The wing colour is grayish-brown with light stripes and spots ( Figs 14–15 ). There are two costal cross veins. Longitudinal vein Rs forked, M unforked in both wings. In hind wing Rs branching from the R very near to the base. Two Rs-M cross veins in both wings. Cua, Cup and Aa veins are very long ones, and the two last longitudinal veins may be regarded as Ja and Jp with high probability ( Fig. 16 ). It is worth mentioning that the number of the cross veins are somewhat larger than in other coniopterygids.

Legs are long and slender. Fourth tarsal segment with lateral lobes, as it is typical in the subfamily Brucheiserinae . First tarsal segment almost as long as the others together. The structure and situation of the plicatures are also the same as in the genus Brucheiser detailed above, i.e., those are in two lateral rows and in a median row on the abdominal segments 3–6 and 2–6 respectively in females, and on 3–7 (lateral) and 2–7 (median) in males, with a glandular structure in the first median plicature.

In male terminalia ( Figs17–19 , 26–29 ) both anterior and posterior appendages of the ectoproct are well separated, and the tenth gonocoxal complex, with or without a ventro-caudal projection is situated above the caudal end of the slender parameres. Anterior appendage forked. Appendages and caudal projection of ectoproct wel sclerotized. (For homologization of the different part of the male terminalia, see later.)

In female terminalia ( Figs 20–23 ) the fused eighth and ninth sternites form a large, strong sclerite. At the base of this sclerite there is a folded glandular structure laterally. Caudal projection of ectoproct well sclerotized. The two gonapophyses laterales fused into a transverse ridge.

In the female internal genitalia ( Figs 24–25 ) the bursa copulatrix large, with two ventro-lateral lobes. Its dorsal and lateral surface covered by slashed projections. Receptaculum seminis weakly sclerotized, sack like. Duct of receptaculum semininis short. Ductus seminalis long, its proximal end connected to the duct of receptaculum seminis, otherwise, its rather wide proximal part is attached to the bursa copulatrix between its lobes. Distal part of ductus seminalis extremely long, thin, looped and coiled. A separate postbursal accessory gland absent.

Remarks: Apart from the striking difference between the wings, the new genus is very close to Brucheiser , the other known genus of the subfamily. The main distinctive features of Flintoconis gen. n. are:

– elongated wings, with very long Cua, Cup and Aa, and with existing J veins;

– presence of the tenth gonocoxal complex instead of penis with U, or horseshoe-shaped sclerotization;

– fused gonapophyses laterales in female terminalia.

Etymology – I dedicate this new genus to Dr. OLIVER FLINT, the excellent American entomologist, who recognized that these unusual insects collected partly by him are coniopterygids.

Figures

Figs 11–13. Flictoconis spp.: 11 = F. gozmanyi sp. n., female paratype, habitus, 12 = Flintoconis sp., female specimen from Aconcagua (Chile), head, 13 = Flintoconis gozmanyi sp. n., female paratype, prothorax, dorsal view. Scale in Fig. 12: 0.3 mm, in Fig. 13: 0.25 mm.

Figs 14–16. Flintoconis gozmanyi sp. n., holotype: 14 = fore wing, 15 = hind wing, 16 = wing venation. Scale: 2.1 mm

Figs 17–18. Male terminalia of Flintoconis gozmanyi sp. n.: 17 = lateral view, 18 = ventral view.IX = 9th sternite, aae = anterior appendage of ectoproct, cpe = caudal projection of ectoproct, dpe = dorsal projection of ectoproct, e = ectoproct, nga = ninth gonapophysis, pa = paramere, pae = posterior ap- pendage of ectoproct, tgc = tenth gonocoxal complex. Scale: 0.15 mm

Fig. 19. Male terminalia of Flintoconis gozmanyi sp. n.: IX = 9th sternite, aae = anterior appendage of ectoproct, cpe = caudal projection of ectoproct, dpe = dorsal projection of ectoproct, nga = ninth gonapophysis, pa = paramere, pae = posterior appendage of ectoproct, tgc = tenth gonocoxal com- plex. Scale: 0.15 mm

Figs 26–27. Flintoconis petorcana sp. n.: 26 = male terminalia, lateral view, 27 = male terminalia, ventral view. IX = 9th sternite, aae = anterior appendage of ectoproct, cpe = caudal projection of ectoproct, dpe = dorsal projection of ectoproct, e = ectoproct, nga = ninth gonapophysis, pa = paramere, pae = posterior appendage of ectoproct, tgc = tenth gonocoxal complex. Scale: 0.3 mm

Figs 28–29. Flintoconis petorcana sp. n.: 28 = male terminalia, caudal view, 29 = tenth gonocoxal complex, ventral view. IX = 9th sternite, aae = anterior appendage of ectoproct, cpe = caudal projection of ectoproct, dpe = dorsal projection of ectoproct, nga = ninth gonapophysis, pae = posterior ap- pendage of ectoproct,tgc = tenth gonocoxal complex.Scale in Fig.28: 0.3 mm, in Fig.29: 0.15 mm

Figs 20–23. Female terminalia and internal genitalia of Flintoconis gozmanyi sp. n.: 20 = female terminalia of a specimen with twirled edges of the fused 8th and 9th sternites, lateral view, 21 = fused 8th and 9th sternites of an other female specimen in expanded position, 22 = female terminalia, ventral view, 23 = female terminalia,caudal view. VIII+IX = the fused 8th and 9th sternites, cpe = caudal projection of ectoproct, dpe = dorsal projection of ectoproct, fgs = folded glandular structure, gl = gonapophysis lateralis, lpl = lateral plicature, mpl = median plicature. Scale: 0.3 mm

Figs 24–25. Female terminalia and internal genitalia of Flintoconis gozmanyi sp. n.: 24 = female internal genitalia, lateral view, 25 = female internal genitalia (without the coiled part of ductus seminalis), dorsal view. bc = bursa copulatrix, ds = ductus seminalis, mo = median oviduct, rs = receptaculum seminis, v = vagina. Scale: 0.15 mm