Phanerotoma macrodentata, Luo & Achterberg & He & Tang & Chen, 2025

Luo, Wen-Juan, Achterberg, Cornelis Van, He, Jun-Hua, Tang, Pu & Chen, Xue-Xin, 2025, Revision of the genus Phanerotoma Wesmael (Hymenoptera, Braconidae, Cheloninae) from China, with descriptions of twenty-four new species, European Journal of Taxonomy 1003, pp. 1-110 : 43-46

Phanerotoma macrodentata sp. nov.

urn:lsid:zoobank.org:act:450468C1-BB80-445E-A8BA-4F4C9E7F4A61

Fig. 17

Diagnosis

Frons punctate-rugose and coriaceous ( Fig. 17C ); head in lateral view protruding dorsally and ocelli invisible ( Fig. 17D ); two apical antennal segments of female much longer than six subapical segments ( Fig. 17K ); vein 1-R1 of fore wing 1.5× as long as pterostigma ( Fig. 17G ), vein 1-CU1 0.6× as long as vein 2-CU1; hypopygium acute apically, lacking an up-curved triangle or apical spine ( Fig. 17I ).

Etymology

Named after the long lower tooth of the mandible, from the Greek ‘ makros ’ for ‘long’ and the Latin ‘ dens ’ for ‘tooth’.

Type material

Holotype

CHINA – Guangdong • ♀; Guangzhou ; Jul. 1982; ZJUH No. 940156a.

Paratypes (8 ♀♀, 38 ♂♂)

CHINA – Guangdong • 1 ♀; same data as for holotype; ZJUH No. 940156b • 2 ♂♂; same data as for preceding; ZJUH Nos 9310903a, 9310903b • 5 ♂♂; same data as for preceding; Sep.–Oct. 1985; Jianshu Li leg.; ZJUH Nos 860657a to 860657e • 5 ♂♂; Shenzhen ; Oct. 1983; Shengzhen Zhou leg.; ZJUH Nos 911090a to 911090e • 13 ♂♂; same data as for preceding; Sep.–Oct. 1985; Jianshu Li leg.; ZJUH Nos 860653a to 860653m • 1 ♀; same data as for preceding; ZJUH No. 860653n . – Fujian • 13 ♂♂; Zhangzhou ; 5 Sep. 1984; Huangquan Wu leg.; ZJUH Nos 881412a to 881412m • 6 ♀♀; same data as for preceding; ZJUH Nos 881412n to 881412s .

Description

Female

MEASUREMENTS. Length of body 3.2 mm, of fore wing 3.0 mm.

COLOURATION. Yellow; stemmaticum dark brown; frons yellowish brown; mandible apically and ovipositor sheath brown; legs pale-yellow.

HEAD. Width 1.7× median length in anterior view ( Fig. 17E ) and part of head above eye in lateral view 0.1 × height of eye ( Fig. 17D ); antenna with 23 segments and 1.2 × as long as fore wing, six subapical antennal segments shortened and cylindrical ( Fig. 17K ), both apical segments elongate, third, fourth and penultimate segments 1.4, 1.5 and 1.7 × as long as wide in lateral view, respectively; area of stemmaticum mainly granulate; OOL: diameter of posterior ocellus: POL=36: 9: 5; eye 1.3 × as long as temple in dorsal view ( Fig. 17C ); frons coriaceous and punctate-rugose, slightly depressed, without median carina; vertex granulate and dull; temple granulate matt and coriaceous; face granulate and with distinct median bump ( Fig. 17E ); clypeus granulate, matt and 0.7× minimum width of face, intertentorial distance 1.7 × minimum width between clypeus and eye, clypeus long erect setose and with 3 indistinct teeth medio-ventrally ( Fig. 17E ); eye large and in lateral view 1.8 × as wide as temple ( Fig. 17D ), in anterior view its height 0.6 × minimum width of face; upper condyle of mandible below lower level of eyes ( Fig. 17E ); malar space 0.9 × as long as basal width of mandible; lower tooth of mandible as long as apical tooth, robust ( Fig. 17L ).

MESOSOMA ( Fig. 17H ). Length 1.8 × its width in lateral view; side of pronotum mainly rugose, but dorsally nearly smooth; mesoscutum densely granulate-rugose and matt, densely setose; notauli hardly indicated; scutellar sulcus wide and with eight carinae ( Fig. 17H ); scutellum triangular, flat, distinctly granulate and matt; metanotum with short median carina anteriorly and small tooth protruding posteriorly; propodeum coarsely reticulate-rugose, with transverse carina, no median carina and latero-posteriorly tuberculate.

WINGS. Fore wing 3.0× as long as its maximum width; length of 1-R1 1.5× as long as pterostigma; r issued relatively close to middle of pterostigma and 0.8× 3-SR; distance between 1-R1 and wing apex 0.3× 1-R1; 2-SR nearly straight ( Fig. 17G ); SR1 straight; 2-SR+M present; parastigma medium-sized; 1-CU1 0.6 × as long as vein 2-CU1, cu-a inclivous and 0.7× as long as 1-CU1; r:3-SR: SR1= 5: 7: 33; 2-SR: 3-SR: r-m=14: 7: 6; r-m reclivous; 2-M weakly curved ( Fig. 17G ).

LEGS ( Fig. 17F ). Hind femur moderately wide and with satin sheen, 4.0× as long as wide and rather widened submedially; hind tibia rather swollen ( Fig. 17A ); middle tibia with medium-sized ivory blister; hind coxa coriaceous and matt.

METASOMA. ( Fig. 17I–J ). Carapace elliptical in dorsal view, 1.9 × as long as wide and as long as mesosoma; first and second tergites coarsely and densely longitudinally rugose; second suture wide; third tergite 1.4× as long as second tergite and laterally weakly curved, in lateral view slightly convex anteriorly, in dorsal view convex medially, densely and semi-circularly rugose and rounded ( Fig. 17I ), lateral lamella hyaline and its margin finely serrate ( Fig. 17I ), and medio-apically not emarginate; ovipositor sheath narrow and parallel-sided ( Fig. 17J ), with some long and erect setae; hypopygium apically acute, lacking an up-curved triangle or apical spine ( Fig. 17J ).

VARIATION. Length of fore wing 2.7–3.0 mm; frons flat or slightly depressed; eye 1.3–1.8 × as long as temple in dorsal view.

Male

Very similar to female, but antennal segments longer and 3–7 subapical antennal segments slender and cylindrical ( Fig. 17B ).

Host

Acrocercops cremerella Snellen, 1904 ( Gracillariidae ).

Distribution

China (Fujian, Guangdong).

Figures

Fig. 17. Phanerotoma macrodentata sp. nov. A, C–L. Holotype, ♀ (ZJUH No. 940156a). B. Paratype, ♂ (ZJUH No. 940156b). A. Habitus, lateral view. B. Apical antenna, lateral view. C. Head, dorsal view. D. Head, lateral view. E. Head, anterior view. F. Hind leg. G. Fore wing. H. Mesosoma, dorsal view. I. First third of metasomal tergites, dorsal view. J. Metasoma, lateral view. K. Antenna, lateral view. L. Mandible, ventral view. Scale bars: A–K=1 mm; L=0.2 mm.