Phaulula javanica, Gorochov, 2025

Gorochov, A. V., 2025, TAXONOMY OF THE KATYDIDS (ORTHOPTERA: TETTIGONIIDAE) FROM EAST ASIA AND ADJACENT ISLANDS. COMMUNICATION 17, Far Eastern Entomologist 514, pp. 1-20 : 10-12

Phaulula javanica Gorochov, sp. n.

https://zoobank.org/NomenclaturalActs/ 7B9E5BB9-AB24-483F-A317-25BAFAE1E690

Figs 21–24 , 29–32

MATERIAL. Holotype – ♂, Indonesia: Java I., “ Java. Soerabaja [Surabaya]. Dr. Fisches 1883”, “17819” ( ZIN).

DESCRIPTION. Male (holotype). Body medium-sized for this genus and more or less matte (but lower half of head, legs, wings and last tergite slightly shining). Coloration uniformly yellowish with greenish tinge, one short dark brown longitudinal stripe on dorsal surface of fore tibia between tympana as well as transparent membranes in most part of stridulatory apparatus of right tgmen and in hind wings (but apical parts of these wings semitransparently yellowish). Upper rostral tubercle with anterior half which almost trice as narrow as scape, having slightly flattened and narrowly rounded apex and distinct median groove on dorsum; lower rostral tubercle very small (almost convexity-like) and clearly not reaching previous tubercle; pronotum with barely sinuate anterior edge, widely rounded posterior edge of disc, rounded lateral bents between disc and lateral lobes (these lobes rather high and having obliquely truncated ventral edges and weekly rounded posteroventral edges), as well as moderately narrow and rounded humeral notches ( Fig. 21 ); tegmina long and rather narrow, with all RS branches starting from R and RA ( Figs 22, 23 ), with stridulatory apparatus as in Figs 24 and 32 , and with barely distinct small fold separating widened proximal part of dorsal field from its narrow distal part in right tegmen ( Fig. 24 ); hind wings somewhat longer than tegmina; legs typical of this genus; last tergite with posteromedian process as in Figs 29, 30 ; cercus slightly arcuate, with distal part strongly curved medially and clearly thickened subapically as well as with denticle-like apex directed medially ( Figs 29– 31 ); epiproct almost longitudinally oval, located under aforementioned process of last tergite and directed downwards/forwards ( Fig 29, 31 ); paraprocts small (shorter than epiproct) and roundly lobule-like; genital plate short, narrowing to rather widely notched apical part (this apical notch moderately deep, almost angular) and with lobules around this notch rather short and moderately thin (almost finger-like but somewhat conical; Figs 29, 31 ).

Female unknown.

Length (in mm). Body 21.5; body with wings 40; pronotum 5; tegmina 31; hind femora 18.3.

The new species belongs to Ph. laevis group and has the male abdominal apex most similar to that of the male from Japan determined by Ichikawa (2004) as “ Ph. macilenta ” (= Ph. gracilis ) and distinguished from Ph. javanica sp. n. by a slightly shining body with fine stripes on the pronotum and tegmina as well as with uniformly greenish fore tibiae (the new species has a mostly matte body and a dark mark on each fore tibia, and its pronotum and tegmina are without any stripes); however, Ph. gracilis was described from Taiwan I. and in accordance to some other authors ( Kang et al., 2014) has this abdominal apex clearly different: the last tergite is with the posterior process very strongly curved downwards ( Figs 42, 43 ). From Ph. apicalis (from Hainan I.) also having a similar male abdominal apex, the new species is distinguished by the narrowest part of the posterior process of the male last tergite somewhat longer and narrower as well as with the subapical part barely lower (see in profile) but having more distinctly angular ventral tubercle near apex (compare Figs 29, 30 and 38, 39 ); from Ph. daitoensis (from Japan), by the apical part of this process not widened and not curved upwards at the apex (compare Figs 29, 30 and 33, 34 ), and clearly narrower tegmina (but for this species, the above-mentioned authors in the same publications indicated somewhat different male genital plates: Kang et al., approximately as in the new species; Ichikawa, with a distinctly wider and less deep posteromedian notch); from Ph. malayica (from Durian I. near Singapore) and Ph. trukkensis (from Caroline Islands), by the male last tergite with a shorter narrowed part of the posterior process, and additionally by this process somewhat higher in the middle part (from Ph. malayica ) and not truncated apically (from Ph. trukkensis ) (compare Figs 29, 30 and 35, 36, 37 ); from Ph. laevis (from Philippines) and Ph. leefmansi (from Karakelong I. between Mindanao and Halmahera), by this process less curved downwards and less convex dorsally (compare Figs 29, 30 and 40, 41, 44, 45 ); and from Ph. dammermani (from Krakatau Islands between Sumatra and Java), by the absence of a pair of distinct dorsal tubercles on the same process. It is also possible that some of these species (including the new one) are only a few subspecies of the same widely distributed species. From all other true and possible congeners, the new species differs in a clearly different shape of the male abdominal apex and/or uniformly light coloration with only a distinct dark mark on the dorsoproximal part of each fore tibia.

ETYMOLOGY. The new species is named after the Java Island where it was collected.

Figures

Figs 21–28. Phaulula, male: 21–24 – Ph. javanica sp. n.; 25–28 – Ph. tapienomima sp. n. Head with pronotum from above/side and in front (21, 26), as well as from side (25); left tegmen (22, 27); distal part of right tegmen in reversed view (23); tegminal stridulatory apparatus from above (24, 28).

Figs 29–48. Phaulula, male: 29–32 – Ph. javanica sp. n.; 33, 34 – Ph. daitoensis; 35, 36 – Ph. malayica; 37 – Ph. trukkensis; 38, 39 – Ph. apicalis; 40, 41 – Ph. laevis; 42, 43 – Ph. gracilis; 44, 45 – Ph. leefmansi; 46–48 – Ph. tapienomima sp. n. Abdominal apex from side (29, 33, 35, 38, 40, 42, 44, 46), from above (30, 34, 36, 37, 39, 41, 43, 45, 47) and from below (31, 48). [33, 34, 38, 39, 42, 43 – after Kang et al. (2014); 35, 36, 40, 41, 44, 45 – after Karny (1926); 37 – after Willemse (1951), modified.]