Sagina apetala Ard., Animadv. Bot. Spec. Alt.: 22. 1764

Dillenberger, Markus S. & Kadereit, Joachim W., 2022, The distinction between Sagina apetala and S. micropetala (Caryophyllaceae: Sagineae), their phylogenetic relationships, and a note on the coastal origin of some widespread ruderals, Willdenowia 52 (1), pp. 5-23 : 17-18

1. Sagina apetala Ard., Animadv. Bot. Spec. Alt.: 22. 1764

≡ Sagina inconspicua Rossmann in Ber. Oberhess. Ges. Natur- Heilk. 8: 24. 1860, nom. illeg. superfl.

≡ Sagina quaternella Schloss. in Schlosser & Vukotinović, Fl. Croat.: 343. 1869, nom. illeg. superfl.

– Lectotype (designated by Crow 1978: 73): Herb. Linn. No. 177.2 ( LINN) . – Note: Linnaeus (1771) did not publish a new name but cited Arduino’s name. Nevertheless, Linnaeus is often incorrectly cited as author of the name. Rossmann (1860) considered the epithet “ apetala ” as meaningless for a plant with (minute) petals and provided a nomenclaturally superfluous (and hence illegitimate) replacement name for S. apetala . Schlosser & Vukotinović (1869) cited “ Moenchia quaternella Alsch. ” as a synonym of S. quaternella . However, they were not citing a basionym but rather what they evidently regarded as a misapplied name – M. quaternella sensu Alschinger , non Ehrh. – because on p. 356 “ Moenchia quaternella Ehrh. ” was cited as a synonym of M. erecta (L.) G. Gaertn. & al. Sagina quaternella was nomenclaturally superfluous when published, and is an illegitimate replacement name for S. apetala , because the latter name was also cited in its synonymy.

= Sagina ciliata Fr., Utkast Sv. Fl. , ed. 3: 713. 1816 ≡ Sagina apetala subsp. ciliata (Fr.) Hook. f., Student Fl. Brit. Isl. : 61. 1870. – Type: not designated. – Protologue: “V. på åkerfält vid Nebelöf nära Ystad”.

= Sagina depressa Schultz, Prodr. Fl. Starg. Suppl. 1: 10. 1819. – Type: not designated. – Protologue: “prope Neobrandenb. auf dem Broda’schen Felde gegen Belvedere, Jun. Jul.”.

= Sagina patula Jord., Observ. Pl. Nouv. 1: 23. 1846 ≡ Sagina apetala var. patula (Jord.) P. D. Sell, Fl. Gr. Brit. Ireland 1: 686. 2018. – Lectotype (designated here): [ France], dans les terres argileuses à Quincieux (Rhône), 2 Jun 1845, A. Jordan s.n. ( LY 0826462 ). – Protologue: “Je l’ai récoltée dans les champs cultivés, à sol argileux, à Quincieux (Rhône)”.

= Sagina filicaulis Jord., Observ. Pl. Nouv. 7: 16. 1849 ≡ Sagina apetala subsp. filicaulis (Jord.) Sennen & Mauricio, Cat. Fl. Rif Orient.: 21. 1933. – Lectotype (designated here): [ France], Toulouse à Blagnac , s.d., Timbal-Lagrave s.n. ( LY 0826452 ). – Protologue: “Je l’ai récoltée à Bedous, dans la vallée d’Àspe, en juillet 1838, et l’ai recue depuis de Toulouse de M. Timbal-Lagrave, d’Angers de M. Boreau, etc.”.

= Sagina lamyi F. W. Schultz in Jahresber. Pollichia 8: 30. 1850 ≡ Sagina apetala subsp. lamyi (F. W. Schultz) Rouy & Foucaud in Rouy, Fl. France 3: 288. 1896. – Lectotype (designated here): [ France], (Cher) Bois du Loire près Saint Florent , May 1848, s.coll. s.n. ( P 05052247 ). – Protologue: “Cher, Saint-Florent, sable du Cher (Tourangin). — Haute-Vien. Magnc-Bourg, roches de Serpentine (Lamy)”. – Note: Schultz based S. lamyi on S. depressa sensu Boreau (1849) , non Schultz. The lectotype was originally in the herbarium of Alfred Déséglise, who was a student of Gustave Tourangin ( Briquet 1940: 206–209). The lectotype is now in P.

= Sagina ambigua J. Lloyd, Fl. Ouest France: 74. 1854. – Type: not designated. – Protologue: ”c. golfe du Morbihan, murs, lieux secs; je l’ai revu à Cadeuil (char.-Inf.), sur les côteaux du Gouessant près Lamballe (C.-Nord), et M. Pontarlier me l’a donné de Challans (Vend.) localités qui sont supposer que cette espece croît sur plusieurs points intermédiaires”.

= Sagina reuteri Boiss., Diagn. Pl. Orient., ser. 2, 1: 82. 1854 ≡ Sagina apetala subsp. reuteri (Boiss.) P. Fourn., Quatre Fl. France: 311. 1936. – Lectotype (designated here): [ Spain], circa Matritum, 1841, Reuter s.n. ( G 00226953 ; isolectotype: G 00226954 ). – Protologue: “Hab. circà Matritum ubi legit vere 1841 amic. Reuter”.

= Sagina urbica Phil. in Linnaea 28: 613. 1857 ≡ Sagina apetala var. urbica (Phil.) Reiche, Fl. Chile 1: 186. 1896. – Lectotype (designated here): [ Chile], Curacasi, Sep 1853, Philippi s.n. (SGO 000001986). – Protologue: “In plateis urbis Santiago frequens, ad Quillota, Curacasi etc.”. – Note: Other original material from Santiago is available at HAL (HAL 0117866).

= Sagina valdiviana Phil. in Linnaea 33: 22. 1864 ≡ Sagina apetala var. valdiviana (Phil.) Reiche, Fl. Chile 1: 186. 1896. – Lectotype (designated here): Chile, Corral, s.d., H. Krause s.n. ( SGO 000001988 ). – Protologue: “ Propo Corral legit orn. Krause”.

= Sagina apetala [unranked or forma] capillaris Lange in Vidensk. Meddel. Naturhist. Foren. Kjøbenhavn, ser. 2, 7: 104. 1865 ≡ Sagina capillaris (Lange) Vayr. in Anales Soc. Esp. Hist. Nat. 30: 508. 1902. – Type: not designated. – Protologue: “In muris Urbis Santiago de Compostela, Aug. c. fl. et fr.”.

= Sagina melitensis Gulia ex Duthie in J. Bot. 13: 37. 1875. – Lectotype (designated by Crow 1978: 73): [ Malta], Insula Melita , Corradino, 13 Mar 1874, Duthie s.n. ( K 000723212 ).

= Alsinella ciliata Greene, Fl. Fran. 126. 1891 ≡ Sagina ciliata (Greene) A. Heller in Muhlenbergia 1: 50. 1904, nom. illeg. [non Sagina ciliata Fr., Utkast Sv. Fl. , ed. 3: 713. 1816] ≡ Sagina ciliata (Greene) Piper in Contr. U.S. Natl. Herb. 11: 259. 1906, isonym ≡ Sagina apetala subsp. ciliata (Greene) A. E. Murray in Kalmia 12: 24. 1982. – Lectotype (designated here): U.S.A., California (Amador Co.), near Ione, Jun 1889, E. L. Greene s.n. (NDG 16653). – Protologue: “Vicinity of Ione”.

= Sagina pachyrrhiza Phil. in Anales Univ. Chile 81: 773. 1892. – Lectotype (designated here): Chile, Santiago , 1888, J. Philippi s.n. ( SGO 000001984 ; isolectotype: SGO 000001985 ). – Protologue : “In plateis urbis Santiago a nepote Julio vere 1888 reperta”.

= Sagina brevipedicellata Sennen in Bol. Soc. Iber. Ci. Nat. 25(9–10): 216. 1927 ≡ Sagina apetala var. brevipedicellata (Sennen) O. Bolòs & Vigo, Butl. Inst. Catalana Hist. Nat., Secc. Bot. 38(1): 86. 1974. – Lectotype (designated here): Spain, Baléares: Pont d’Inca, lieux sablonneux, 9 Apr 1919, F. Bianor s.n. ( BC 11053 ; isolectotype: P 05398131 ). – Protologue: “Hab. Baléares: Pont d’Inca, lieux sablonneux. Leg. F. Bianor”.

= Sagina rodriguezii var. condensata Sennen. Diagn. Nouv. : 144. 1936. – Lectotype (designated here): Spain, Barcelona: Premia, bords de la Méditerranée, 29 Mar 1932, F. Sennen 8213 ( BC 75090 ). – Protologue: “Hab.—Barcelona: Premia de Mar , étroite bande entre la voire ferrée et la mer”. – Note: “ Sagina condensata ” was cited by Sennen in the synonymy of S. rodriguezii var. condensata (l.c.) and is therefore not validly published.

= Sagina vasconica Sennen, Diagn. Nouv. : 145. 1936. – Lectotype (designated here): Spain, Alava : Pipahon, montagne, 800 m, 1932, M. Losa s.n. ( BC 75091 ; isolectotype: MPU 021876 ). – Protologue: “Hab.— Alava: Pipahon, montagne, 800 m. Leg. Dr. M. Losa”.