Anemesia gidzhovak Zamani and Fomichev, 2025

Zamani, Alireza & Fomichev, Alexander A., 2025, Four new species of Mygalomorphae (Araneae) from Tajikistan and Afghanistan, Journal of Natural History 59 (13 - 16), pp. 775-793 : 776-781

Anemesia gidzhovak Zamani and Fomichev sp. n.

( Figures 1–18 , 45 , 47–48 )

Types

♂ holotype ( ISEA, 001.9166) and 1♀ paratype ( ISEA, 001.9167); TAJIKISTAN: Khatlon Region, Vakhsh Mt . Range, Gidzhovak Village , 38.1826°N, 69.3674°E, 1200 m, 2 May 2024, leg . A .A. Fomichev.

Etymology

The specific epithet, a noun in apposition, refers to the type locality of the species.

Diagnosis

The new species belongs to the incana group sensu Zonstein (2018b), which currently comprises three species: Anemesia incana Zonstein, 2001 ; A. infumata Zonstein, 2018 ; and A. infuscata Zonstein, 2018 . The male of A. gidzhovak sp. n. differs from those of the aforementioned three species by having a longer embolus that gently curves in its distal part, and the ratio of the embolus length to the width of the bulbal part of the palpal organ 1.7, whereas it is 1.3 in A. incana and A. infuscata , and 1.5 in A. infumata . Moreover, the embolus in the other three species is less distinctly bent ( Figure 13 , cf. Zonstein 2018b, figs 215–223).

The female differs from that of A. infumata by having distinctly globular spermathecal heads that are clearly wider than the corresponding spermathecal stalks ( Figure 18 , cf. Zonstein 2018b, figs 255–257), from that of A. infuscata by having shorter, almost columnar stalks (these being noticeably longer and more strongly narrowing apically in A. infuscata ; Figure 18 , cf. Zonstein 2018b, figs 258–265), and from that of A. incana by having less defined spermathecal heads (these being more developed and much wider than corresponding stalks in A. incana ; Figure 18 , cf. Zonstein 2018b, figs 251–254).

Description

Male. Habitus as in Figure 2 . Total length 16.5. Carapace 7.0 long, 6.1 wide. Sternum 3.3 long, 3.0 wide. Labium 0.6 long, 1.1 wide. Eye sizes and interdistances ( Figure 3 ): AME 0.20 (0.33), ALE 0.36, PLE 0.26, PME 0.17, AME–AME 0.21 (0.11), ALE–AME 0.13 (0.10), ALE–PLE 0.11, PLE–PME 0.07, PME–PME 0.57. Cheliceral furrow with 8 pro- and 7 retromarginal teeth ( Figure 7 ). Each maxilla with ca. 30–40 cuspules. Carapace, chelicerae, and labium brown. Maxillae, sternum, and leg coxae light brown ( Figure 9 ). Palp: Fe–Pa brown, Ti–Ta light brown. Leg I: Fe–Ti brown, Mt–Ta orange-coloured. Legs II–IV: Fe–Pa brown, Ti light brown, Mt–Ta yellow. Dorsal side of abdomen yellow-grey, with longitudinal dark stripe medially and dark spots laterally. Ventral side of abdomen and book-lung covers yellow. Spinnerets pale yellow. Spination of palp and legs: palp: Fe d3 p1, Pa p1, Ti p5 r4 v5; leg I: Fe d5 p4 r4, Pa p1, Ti p5 r4 v7, Mt d1 p3 r1 v6; leg II: Fe d4 p3 r2, Pa p2, Ti p5 v8, Mt p3 r2 v7; leg III: Fe d6 p3 r4, Pa d1 p3 r2, Ti d3 p6 r3 v6, Mt d3 p6 r3 v8, Ta p1; leg IV: Fe d4 p3 r3, Pa p1, Ti d2 p7 r4 v9, Mt d2 p6 r3 v11, Ta p2. Distal megaspine on tibia I 1.5× longer than metatarsus I wide. Measurements of palp and legs: palp: 11.85 (4.5, 2.35, 3.75, -, 1.25); leg I: 24.35 (7.15, 3.5, 5.4, 5.35, 2.95); leg II: 21.95 (6.55, 3.0, 4.65, 4.75, 3.0); leg III: 19.95 (5.6, 2.5, 3.6, 5.0, 3.25); leg IV: 25.15 (6.75, 2.5, 5.65, 6.75, 3.5). Spinnerets ( Figure 5 ): PMS 0.5 long, 0.28 in diameter; PLS maximal diameter 0.68, length of its basal, medial and apical segments 1.25, 0.63, 0.58, respectively (total length 2.46).

Palp as in Figures 11–16 ; bulbal part of palpal organ asymmetrically oval; embolus long and tapering, with a gentle but distinct curvature in its distal part.

Female. Habitus as in Figure 1 . Total length 22.3. Carapace 7.8 long, 6.7 wide. Sternum 4.0 long, 4.2 wide. Labium 1.25 long, 2.0 wide. Eye sizes and interdistances ( Figure 4 ): AME 0.19 (0.27), ALE 0.39, PLE 0.29, PME 0.17, AME–AME 0.24 (0.11), ALE– AME 0.24 (0.20), ALE–PLE 0.16, PLE–PME 0.09, PME–PME 0.56. Cheliceral furrow with 7 pro- and 9 retromarginal teeth ( Figure 8 ). Each maxilla with ca. 20–30 cuspules. Carapace, chelicerae, sternum, labium, maxillae, and leg coxae brown ( Figure 10 ). Palps and legs brown on proximal segments, paler on distal segments. Abdomen dirty yellow, with indistinct grey dorsal pattern. Book-lung covers and spinnerets yellow. Spination of palp and legs: palp: Fe p1, Pa d1, Ti p3 v10, Ta p1 v2; leg I: Fe p1, Ti v4, Mt v6; leg II: Fe p1, Pa p1, Ti p2 v4, Mt p1 v7; leg III: Pa p3, Ti d1 p2 r1 v3, Mt d1 p7 r3 v9, Ta p5; leg IV: Ti r2 v4, Mt p2 r2 v15, Ta p5 v1. Measurements of palp and legs: palp: 12.65 (4.5, 2.4, 3.0, -, 2.75); leg I: 19.6 (6.2, 3.55, 4.05, 3.4, 2.4); leg II: 17.45 (5.4, 3.25, 3.45, 3.1, 2.25); leg III: 14.8 (4.45, 2.65, 2.5, 2.95, 2.25); leg IV: 20.25 (5.8, 3.0, 4.45, 4.3, 2.7). Spinnerets: PMS 0.75 long, 0.5 in diameter; PLS maximal diameter 1.13, length of basal, medial and apical segments 1.25, 0.63, 0.75, respectively (total length 2.63).

Spermathecae as in Figures 17–18 ; stalks stout, only slightly narrowing apically; heads broadly oval; pore glands present on both parts.

Habitat

Grassland on loess hills ( Figure 45 ).

Distribution

Known only from the type locality in the Khatlon Region of south-western Tajikistan ( Figure 47 ).

Figures

Figures 1–5. Habitus (1–2), eye tubercle (3–4), and spinnerets (5) of male (2–3, 5) and female (1, 4) of Anemesia gidzhovak sp. n. 1–4 = dorsal view; 5 = lateral view. Scale bars:1–2 = 5 mm; 3–4 = 0.2 mm; 5 = 1 mm.

Figures 6–10. Tibia and metatarsus of left leg I (6), left chelicera (7–8), and cephalothorax (9–10) of male (6–7, 9) and female (8, 10) of Anemesia gidzhovak sp. n.; 6 = retrolateral view; 7–10 = ventral view. Scale bars: 6–8 = 1 mm; 9–10 = 2 mm.

Figures 11–16. Left palp (11–12) and palpal organ (13–16) of Anemesia gidzhovak sp. n.; 11, 14 = prolateral view; 12, 16 = retrolateral view; 13 = anterior view; 15 = posterior view. Scale bars: 11–12 = 1 mm; 13–16 = 0.2 mm.

Figures 17–18. Spermathecae of Anemesia gidzhovak sp. n., dorsal view; 17 = both spermathecae; 18 = left spermatheca. Scale bars = 0.2 mm.

Figures 43–46. Living male of Ummidia dudkoi sp. n. (43), and habitat at type localities of Raveniola farkhor sp. n. and U. dudkoi sp. n. (44), of Anemesia gidzhovak sp. n. (45), and of Raveniola zonsteini sp. n. (46). Photographs: A.A. Fomichev (43–45) and C. Reuter (46).

Figures 47, 48. Type localities of the new mygalomorph species. Diamond = Anemesia gidzhovak sp. n.; triangle = Raveniola zonsteini sp. n.; square = Raveniola farkhor sp. n. and Ummidia dudkoi sp. n. Figure 47 corresponds to the inside of the frame in Figure 48. National borders are shown in black, rivers and lakes are shown in blue.