Clupeonella Kessler, 1877

Eagderi, Soheil, Mouludi-Saleh, Atta, Esmaeili, Hamid Reza, Sayyadzadeh, Golnaz & Nasri, Manoochehr, 2022, Freshwater lamprey and fishes of Iran; a revised and updated annotated checklist- 2022, Turkish Journal of Zoology 46 (6), pp. 500-522 : 505

Genus Clupeonella Kessler, 1877

17. Clupeonella caspia Svetovidov, 1941 [N]-[LC], Caspian tyulka/Caspian Sea Basin ( Esmaeili et al., 2018; Amiri et al., 2018).

Remark: Clupeonella cultriventris is widespread along the coasts of the Black and Caspian Seas, entering the lower reaches of rivers. Based on unpublished data (see Kuljanishvili et al., 2020) including molecular and morphological studies they failed to distinguish the Caspian population ( C. caspia ) from the Black Sea population ( C. cultriventris ). Hoestlandt (1991) followed by Kottelat and Freyhof (2007) separate these two tyulkas largely based on the length of paired fins, a character that is found to be very much overlapping in the materials examined by Kuljanishvili et al. (2020). Therefore, they treated C. caspia as a synonym of C. cultriventris . More data using different populations from the southern Caspian and the Black seas is needed to accept this synonymy.

18. Clupeonella cultriventris (Nordman, 1840) [N]- [LC], Caspian tyulka/Caspian Sea Basin ( Amiri et al., 2017a,b; Esmaeili et al., 2018).

19. Clupeonella engrauliformis (Borodin, 1904) [N]-[NE], Caspian tyulka/Caspian Sea Basin ( Esmaeili et al., 2018).

20. Clupeonella grimmi Kessler, 1877 [N]-[NE], Southern Caspian sprat, Bigeye kilka/Caspian Sea Basin ( Esmaeili et al., 2018).