Germainia khasyana Hack. — Plate, 1891

Teerawatananon, A., Sungkaew, S., Boontia, V. & Hodkinson, T. R., 2013, A taxonomic revision of Germainia (Andropogoneae: Poaceae) in Thailand, Blumea 57 (3), pp. 221-228 : 225

2. Germainia khasyana Hack. — Plate 1

Germainia khasyana Hack.(1891) 50.— Themeda capitata (Balansa & Poitr.) Roberty var. khasyana (Hack.) Roberty (1960) 102, nom. inval. — Type: Clarke 44830B (lecto W, sheet W­Hackel­1916­00027613; iso W, sheet W­Hackel­1916­00027614), India or Khasya, Nartung, 1220 m.

Perennial. Culms slender, 25– 40(–75) cm tall, erect, branched; nodes pilose. Leaf­sheaths 1.5 –3 cm long, pilose with tubercle-based hairs, margins scarious. Ligule an eciliate membrane, 1–1.5 mm long. Leaf­blades lanceolate to linear-lanceolate, 2– 4.5 by 0.3 – 0.4 cm, upper surface and margins pilose with tubercle-based hairs, margins thick. Racemes solitary, capitate, 1–1.7 cm long, composed of (2–)3(–4) sessile and pedicelled spikelets; peduncles slender, 6 –14 cm long, waxy. Sessile spikelets yellowish green, oblong, 10–17 by 3– 4 mm; lower glumes coriaceous, oblong, 6.5–12 by 3 – 4 mm, glabrous, truncate to muticous, upper margin ciliate, 7–9(–11)-nerved, nerves not anastomosing; upper glumes lanceolate to linear-lanceolate, 8 –17 by 2 –3 mm, pubescent on dorsal surface on the upper half, obtuse, upper margin ciliate, 3-nerved; lower lemmas linear-lanceolate, 6 –14 mm long, obtuse, upper margin ciliate, 1-nerved; lower paleas linear-lanceolate, 6 –14 mm long, obtuse, upper margin ciliate; upper lemmas absent; upper paleas similar to the lower ones; stamens 2, anthers purple, 4 –7 mm long. Pedicelled spikelets lanceolate, 6– 6.5 by 0.8–1(–2) mm, caducous; pedicels 2.5– 4 mm long, sparsely hispid; spikelet callus linear, c. 1.5 mm long, attached obliquely, hairy, hairs 1– 2 mm long; lower glumes lanceolate to ovate-lanceolate, 6 – 6.5 by 1–2 mm, pubescent, truncate or obtuse, upper margin ciliate, 5 –7-nerved; upper glumes lanceolate to ovate-lanceolate, 5– 5.5 by 1–1.8 mm, pubescent, truncate or obtuse, upper margin ciliate, 3-nerved; lower florets suppressed; upper lemmas linear, 2.5– 4 mm long, awns 3.5 –5.5 cm long, columns 2 – 4 cm long; upper paleas oblong, 2.5 –3 mm long, truncate and dentate, upper margin ciliate; styles 2, stigmas white, 5–7 mm long. Caryopsis oblong, 3– 3.2 mm long.

Distribution — Thailand (N: Phitsanulok). This species is also found in India (Meghalaya) and Myanmar (Kachin).

Habitat & Ecology — Open swampy areas, tropical grasslands, 150 –1200 m altitude.

Typification notes — Hackel (1891) typified G. khasyana by citing the specimens of Clarke 44830 and Clarke 42558. Roberty (1960: 102) selected the specimen of Clarke 14568 as the type of G. khasyana . However that collection was not cited by Hackel (1891) and was not found in his herbarium (W) (J.F. Veldkamp, pers. comm.). Later, Chai-Anan (1972: 41) designated Clarke 44830A (BM, K) as hololectotypes, but this lectotypification is incomplete as two sheets were mentioned. We have found that there are five sheets of Clarke 44830: Clarke 44830A (K), Clarke 44830B (W, 2 sheets), Clarke 44830C (K) and Clarke 44830D (BM) and three sheets of Clarke 42558: Clarke 42558A (K), Clarke 42558D (BM) and Clarke 42558J (RGB). Therefore, there are at least eight specimens of Clarke which could be selected as the holotype. We hereby designate Clarke 44830B as the lectotype. This lectotype is the one housed in his herbarium W (W­Hackel­1916­00027613) and that bears a label with Hackel’s handwriting (J.F. Veldkamp, pers. comm.).

Specimens examined. Clarke 6447, 42558A, 42558D, 42558J, 44830A, 44830C, 44830D; Hansen et al. 11231; Lace 6054; Teerawatananon & Sungkaew 906, 907.

Figures

Plate 1 a, b. Germainia capitata Balansa & Poitr.a. Habit;b. inflorescence.— c, d. Germainia khasyana Hack. c. Habit; d. inflorescence (a, b: Teerawatananon & Sungkaew 834; c, d: Teerawatananon & Sungkaew 906). — Photos: all A. Teerawatananon.