Madrasostes bartolozzii Ballerio, 2013

Ballerio, Alberto & Maruyama, Munetoshi, 2025, Review of Madrasostes Paulian, 1975 (Coleoptera: Hybosoridae: Ceratocanthinae) from Vietnam, with remarks on its generic limits, Zootaxa 5661 (3), pp. 420-430 : 421

Madrasostes bartolozzii Ballerio, 2013

( Figs. 1 ; 4B )

Madrasostes bartolozzii Ballerio, 2013: 380 (description and iconography).

Type locality. Ba Be National Park (Bac Kan Province) .

Material examined. Holotypus, ♀ (VNMN) and paratypus ♀ (MZUF): Northern Vietnam, Bac Kan Province, Ba Be National Park (~ 350 m), 22°25’0.69”N 105°37’53.16”E, 3-8.VI.2011, L. Bartolozzi, S. Bambi, F. Fabiano & E. Orbach leg. ; 2 ♂♂: Ba Be 220m alt. Cao Bang prov. / [N. Vietnam] 12.V.1998 S. Nomura leg. (ABCB, NSMT) ; 1 specimen of sex unstated: Bo Lu (140 m) Ba Be Cao Bang prov. [N. Vietnam] 13.V.1998 S. Nomura leg. (NSMT) . Complementary description and remarks. The availability of male specimens allows for the first illustrations of the aedeagus (L = 0.8 mm) ( Fig. 1C ) and spiculum gastrale (L = 0.7 mm) ( Fig. 4B ), as well as to document the sexual dimorphism in this species, which was originally described from two females only. Males differ in several features: notably, they lack the acute projection at the apex of the clypeus seen in females, furthermore, as happens in the majority of other Madrasostes , the inner apical spur of the mesotibiae is bent sharply inward at a right angle, and the apical spur of the protibiae is more strongly curved inward. The additional specimens listed here show little variation from the type series. However, the original description requires amendment. Closer examination of the head revealed the presence of a short, narrow genal canthus that does not reach the occipital margin and delimits a very narrow dorsal ocular area ( Fig. 1B ) .

Figures

FIGURE 1. Madrasostes bartolozzii Ballerio, 2013.A. Distended male in dorsal view. B. Detail of pronotum and head, showing the sinuature around fore pronotal angles, and the genal canthus with the small dorsal ocular area. C. Aedeagus in dorsal, lateral and ventral views.

FIGURE 4. Spicula gastrales of A. Madrasostes nomurai Ballerio & Maruyama sp. nov., B. Madrasostes bartolozzii Ballerio, 2013 and C. Madrasostes deharvengi Gao, 2009.