Mucuna sakapipei Wiriad., 2016

Wiriadinata, H., Ohashi, H. & Adema, F., 2016, Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 16. The genus Mucuna, Blumea 61 (2), pp. 90-124 : 117

37. Mucuna sakapipei Wiriad. , sp. nov. — Fig. 7

Twigs glabrous, (thinly) puberulous or very thinly sericeous. Petioles 4–11 cm long. Leaflets ovate,5.5–12.2 by 3–7 cm, acumen 5–11 mm long,nerves 3–6 per side. Inflorescences axillary,pseudoracemes,8–11.5 cm long. Calyx 10–11 mm long. Corolla pale green. Ovules 2. — Type: De Haan 1773 (holo BO; iso A, K, L), Halmahera, Distr. Weda, Pajahi road, 26.11.1950 .

Liana. Twigs terete, striate, diam 1–6 mm, glabrous, (thinly) puberulous or very thinly sericeous. Stipules caducous. Petioles 4 –11 cm long, striate, puberulous or with scattered appressed hairs; rachis mostly as the petiole, 1.5–3.2 cm long; pulvinus 3–6 mm long. Stipellae acicular, 2.4–3.8 by 0.1 mm, glabrous or with some hairs. Leaflets: terminal ovate, 6–12.2 by 3–7 cm, index 1.6–2.1, base rounded or truncate, apex acuminate, acumen 5–11 mm long, above with scattered appressed hairs, below with scattered appressed hairs or thinly sericeous, midrib and nerves slightly raised above, nerves 3–6 per side, 5–46 mm apart, anastomosing near the margin; lateral mostly as the terminal, 5.5–12 by 3–7 cm; pulvinus 3–7 mm long. Inflorescences axillary, pseudoracemes, 8–11.5 cm long, peduncle 7–10 cm long, at base glabrous to sericeous at apex or thinly puberulous at base becoming denser upwards. Bracts to the brachyblasts ovate, 6–7 by 2.5 mm, outside sericeous, inside glabrous. Brachyblasts 1–3 mm long. Bracts to the flowers ovate, c. 7.5 by 3.0 mm, outside sericeous, inside glabrous. Pedicels 10–16 mm long, ferruginous puberulous. Bracteoles caducous. Calyx in bud cup-shaped, at anthesis broadly campanulate, 10–11 mm long, tube c. 6 mm long; upper lip semicircular, 4 by 14 mm, bidentate, lateral teeth triangular, 2–4 by 5 mm, median tooth triangular, 4–5 by 6 mm; both sides (ferruginous) sericeous. Corolla pale green. Standard: claw 2–3 mm long, with some hairs at apex; blade ± orbicular, 2–2.1 by 1–2 cm, outside with some hairs at base, inside glabrous. Wings: claw 5 mm long, ciliate along lower margin; blade elliptic-falcate, 2.7–3 by 0.5–0.7 cm, auricle 3 mm long, outside sericeous at auricle and just above, ciliate along lower margin at base, inside sericeous along lower margin. Keel petals: claw 5–7 mm long; blade falcate, 2.5–2.6 by 0.4 cm, auricle inconspicuous, hard part c. 5 mm long, lateral pocket inconspicuous, outside with some hairs between claw and auricle. Stamens 2.5 cm long, tube 2 cm long, glabrous; anthers of longer filaments 0.1 mm long, bearded, of shorter filaments 0.2 mm long, with some hairs. Disc c. 1.0 mm high. Ovary 4.5– 5 mm long, sericeous; ovules 2; style 22 mm long, sericeous, apical part glabrous. Pods flattened ellipsoid, 6.0–7.5 by 2.5–4.5 cm, reddish brown irritating hairs, upper wing 4–7 mm wide, lower wing 4–8 mm wide, with transverse, 6–8 mm high lamellae, valves with few hairs and irritating hairs. Seeds discoid, 13–16 by 13–15 by 3–3.6 mm; hilum 33 mm long, 4/5 of the circumference.

Distribution — Malesia: Moluccas: Halmahera.

Habitat & Ecology — Primary forest . Soil: volcanic tuff. Altitude up to 20 m. Flowering: November; fruiting: September, November .

Note — The flowers of the only flowering specimen available to Adema at L are insect-damaged. Some of the characters could not be fully described.

Figures

Fig. 7 Mucuna sakapipei Wiriad. a. Piece of twig with a leaf and an inflorescence; b. detail of leaflet lower surface; c. calyx opened, from outside; d. detail of calyx indumentum; e. standard; f. wing petal; g. keel petal; h. ovary and lower part of style; i. upper part of style and stigma; j. infructescence and fruit; k. detail of indumentum and ornamentation; l. seed (all: De Haan 1773). — Drawing Margaret Tebbs.