Cranionycta (Cranionycta) oda transversa ( Kozhanchikov, 1950 ), 2025

Kiss, Ádám, 2025, Taxonomic study of the genus Cranionycta de Lattin, 1949 (Lepidoptera: Noctuidae: Acronictinae) with description of a new subgenus and 21 new species, Zootaxa 5640 (1), pp. 1-71 : 17-18

Cranionycta (Cranionycta) oda transversa ( Kozhanchikov, 1950) stat. rev.

( Figs 17–20 , 63, 64 , 82 , 92, 93 , 117 , 133 )

Craniophora transversa Kozhanchikov, 1950 , Lepidoptera , Orgyiidae, Fauna SSSR 12: 540, fig. 280 (male and female genitalia). Type-locality: Korea. Lectotype: male, in coll. ZISP, here designated.

Lectotype designation. Lectotype of Craniophora transversa Kozhanchikov, 1950 : male, labelled as “ 27/7 ” [handwritten], “typus” [greenish label with cut edges], “ nov. sp? / Corea ” [handwritten], “ Acronycta / coreana nov. / S Tshetverikov / det. ” [handwritten], “Кол. Вел. Кн. / НиколаЯ / Михаиловича” [Kol.[lekcija]. vel.[ikogo] kn.[jaza] / Nikolaya / Mikhailovicha; with thin black line next to the upper, down and right sides of the label], “slide / Matov0629” [handwritten with blue ink]. Lectotype is deposited in the ZISP.

Material examined. [ Korea]. Paralectotype, female, 27.vii., ex. coll. N. Mikhailovicha, slide No.: KA757f ( ZISP) . Korea. 1 male, ex. coll. Staudinger, slide No.: KA 1107m ( MfN) ; Paralectotype, female, 24.vii., ex. coll. Erschov ( ZISP) ; Paralectotype, female, 27.vii., ex. coll. N. Mikhailovicha ( ZISP) ; Paralectotype, female, 4.viii., “praep. micr. No.: 3489”, ex. coll. N. Mikhailovicha, slide No.: Matov 0348 ( ZISP) ; 1 male, Kongosan, Utikongo , 500 m, 27.vii.1940, leg. H. Höne, slide No.: KA 678m ( ZFMK) ; 2 females, Seoul, 6.vii.1959, 9.vii.1959, leg. S.W. Pak, ex. coll. S. Sugi, 29-0057808, 29-0057809, slide No.: SS 909♀ ( NIAES) . North Korea. Prov. South Hwanghae. 1 female, Mt. Suyong-san, Haeju , 31.vii.1982, No. 894, leg. L. Forró & L. Ronkay, KOREA database No.: 2010- 02251, slide No.: KA044f ( HNHM) . South Korea. Prov. Gyeonggi. 1 female, Mt. Hwaya, Gapyeong , 16.vi.2004, 37°40'N, 127°25'E, MPNE0909291025, ex. coll. MNU, slide No.: KA1226f ( HNHM) . Prov. North Chungcheong. 1 female, Mt. Sobaek, Danyang , 21.vii.2005, 37°02'N, 128°31'E, 300 m, leg. S.-W. Choi, MPNE0909291030, ex. coll. MNU, slide No.: KA1227f ( HNHM) . Prov. Kangwon. 1 female, Bongyeong-ri, Exp. Forest Station , 37°46'N, 127°48'E, 230 m, 30.vi.2004, leg. H.L. Han, slide No.: KA1825f ( HNHM) ; 1 female, Chuncheon-dam , 37°57'34.5"N, 127°39'33.1"E, 170 m, 26.viii.2003, No. 1736, leg. A. Kun & M. Földvári, slide No.: KA211f ( HNHM) .

Note. “ Craniophora transversa ” was described by Kozhanchikov (1950) based on an indeterminate number of specimens from Russia, Primorsky Krai (Suchan and Sedanka) and Korea with no correct type designation. In the collection of the Zoological Institute, Saint-Petersburg, there are six old specimens from Suchan and Sedanka (actually belonging to the nominotypical subspecies), and five old specimens from Korea. All Korean specimens bear the label “ Acronycta coreana nov. S. Tshetverikov det.” or “ Acronycta coreana Tshtv. S. Tshetverikov det.”. It is possible that Tshetverikov realized the differences between the Russian Far Eastern and Korean populations, however, he has never described it. In order to justify preserving the name “ transversa ” instead of introducing a new one ( “ coreana ”), a Korean specimen was selected for the lectotype .

Diagnosis. Cranionycta (Cranionycta) oda transversa ( Figs 17–20 ) can be distinguished from the nominotypical subspecies ( Figs 14–16 ) by the generally darker brownish forewing; the less contrast, less conspicuous subterminal line and terminal field (the latter less whitish); the somewhat less pronounced orbicular and reniform stigmata; the more reduced discal spot on the hindwing; from C. (N.) delattini ( Figs 29, 30 ) by the somewhat more cold brownish forewing; the narrower, longer tornal streak; the straighter medial line with more, smaller waves; the less pronounced postmedial line without the narrow black stripe outwardly between the veins R3 and M2; the more conspicuous orbicular stigma with a longer blackish inner outline; and the more conspicuous reniform stigma with a blackish inner half. Wingspan 30–35 mm.

Cranionycta (Cranionycta) oda transversa ( Figs 63, 64 , 82 ) has a proportionally larger, tooth-like cornuti on the mid-lateral surface of the vesica; a less sharp angled and somewhat wider terminal twist of the appendix bursae ( Figs 92, 93 ); and a narrower oval window in the male 8 th tergite ( Fig. 117 ) than in C. (C.) o. oda ( Figs 60–62 , 81 , 90, 91 , 116 ).

Distribution. Cranionycta (Cranionycta) oda transversa occurs only on the Korean Peninsula.

The formosana species-group

This species group contains only one species with several unique features. The external features that diagnose formosana from other Cranionycta species are: 1) the shorter and thinner darker area between the basal and antemedial lines at the costa; 2) the reduced, short, reversed (widest at the postmedial line), somewhat triangular tornal streak; 3) the contrasting antemedial line; 4) the strongly reduced orbicular stigma marked only with a lighter spot or tiny, blackish dots inwardly; 5) the reduced reniform stigma marked with tiny, blackish dots on the veins; and 6) the brownish suffused hindwing with a more pronounced marginal band and postdiscal line. The typical male genital features are 7) the slightly concave, in the basal half slightly sclerotized medial sclerite; and 8) the cornuti patch on the mid-lateral surface of the vesica consisting of several, more or less equal-sized, tiny, sclerotized tooth-like spinules. The female genitalia are characterised by 9) the widest at base, more elongate, distally slightly lobe-like anterior apophysis; and 10) the straighter appendix bursae having a more turned, slenderer terminal twist.

In the male 8 th abdominal segments, the sternite is slightly wider at the distal edge with a wider than long window; tergite is wider in the middle with a less oval window.

In the female 7 th abdominal segments, the sternite has a weaker sclerotization than in the oda species-group.

Distribution. Taiwan.

Figures

FIGURES 11–20.Adults of Cranionycta spp.11. C. (C.) muscosa sp. nov., HT, male, China, Taihang Mts, Beijing, Xiaolongmen Nat. Forest Park, slide No.: KA756m (photo and coll. ZISP); 12. C. (C.) muscosa sp. nov., PT, female, China, Taihang Mts, Beijing, Xiaolongmen Nat. Forest Park, slide No.: KA755f (photo and coll. ZISP); 13. C. (C.) muscosa sp. nov., PT, male, China, Beijing, Xiaolongmen, [abdomen missing] (photo and coll. NEFU); 14. C. (C.) oda oda, HT, male, Russia, Primorsky Krai, Suchansky Rudnik [Partizansk], (photo and coll. ZfBS); 15. C. (C.) oda oda, AT, female, Russia, Primorsky Krai, Suchansky Rudnik [Partizansk], (photo and coll. ZfBS); 16. C. (C.) oda oda, male, China, Prov. Liaoning, Fushun city, Douling, slide No.: HHL6193 (photo and coll. NEFU); 17. C. (C.) oda transversa, LT of Craniophora transversa, male, Korea, slide No.: Matov0629 (photo and coll. ZISP); 18. C. (C.) oda transversa, female, South Korea, Prov. Kangwon, Chuncheon-dam, slide No.: KA211f (coll. HNHM); 19. C. (C.) oda transversa, male, Korea, slide No.: KA1107m (coll. MfN); 20. C. (C.) oda transversa, female, South Korea, Prov. North Chungcheong, Mt. Sobaek, Danyang, slide No.: KA1227f (coll. HNHM).

FIGURES 61–66. Male genitalia of Cranionycta spp. 61. C. (C.) oda oda, Russia, Primorsky Krai, Gornotaezhnaya Station, slide No.: KA863m (valvae) (coll. ZMUC), Razdolnoye, slide No.: KA180m (phallus) (coll. HNHM); 62. C. (C.) oda oda, Russia, Primorsky Krai, Razdolnoye, slide Nos: KA173m (valvae), KA177m (phallus) (coll. HNHM); 63. C. (C.) oda transversa, LT, Korea, slide No.: Matov0629 (photo and coll. ZISP); 64. C. (C.) oda transversa, Korea slide No.: KA1107m (coll. MfN); 65. C. (C.) formosana, HT, Taiwan, Nantou County, Hueisun Forest, MHNG ENTO 60282, slide No.: KA2620m (coll. MHNG); 66. C. (C.) formosana, PT, Taiwan, Nantou County, Hueisun Forest Area, slide No.: KA2492m (photo and coll. GR).

FIGURES 73–83. Male genitalia and magnified part of the vesica of Cranionycta spp. 73. C. (N.) fuscothoracica sp. nov., PT, Vietnam, Plato Tay, Nguyen, Mt. Ngok Linh, slide No.: KA2073m (photo L. Ronkay; coll. MfN); 74. C. (N.) boursini sp. nov., HT, Nepal, Thodung, slide No.: N956 (photo and coll. ZSM); 75. C. (N.) jubingica sp. nov., HT, Nepal, Jubing, slide No.: KA1222m (coll. ZSM); 76. C. (N.) albothoracica sp. nov., PT, Vietnam, Plato Tay, Nguyen, Mt. Ngok Linh, slide No.: KA2072m (photo L. Ronkay; coll. MfN); 77. C. (N.) falcata sp. nov., HT, Nepal, Syangja district, 14 km E of Syangja, slide No.: KA2429m (photo and coll. GR); 78. C. (C.) aurata sp. nov., HT, China, Prov. Sichuan, near Jiuzhaigou, slide No.: GYP2886 (coll. PG); 79. C. (C.) inquieta, China, Prov. Jilin, Changbaishan, slide No.: HHL6187 (photo and coll. NEFU); 80. C. (C.) muscosa sp. nov., HT, China, Taihang Mts, Beijing, Xiaolongmen Nat. Forest Park, slide No.: KA756m (photo and coll. ZISP); 81. C. (C.) oda oda, Russia, Russian Far East, Barabash, slide No.: KA862m (coll. ZMUC); 82. C. (C.) oda transversa, Korea, slide No.: KA1107m (coll. MfN); 83. C. (C.) formosana, PT, Taiwan, Nantou County, Hueisun Forest Area, slide No.: KA2492m (photo and coll. GR).

FIGURES 90–95. Female genitalia of Cranionycta spp. 90. C. (C.) oda oda, Russia, Primorsky Krai, Razdolnoye, slide No.: KA175f (coll. HNHM); 91. C. (C.) oda oda, Russia, Primorsky Krai, Razdolnoye, slide No.: KA179f (coll. HNHM); 92. C. (C.) oda transversa, PLT, Korea, slide No.: slide No.: Matov0348 (photo and coll. ZISP); 93. C. (C.) oda transversa, South Korea, Prov. North Chungcheong, Mt. Sobaek, Danyang, slide No.: KA1227f (coll. HNHM); 94. C. (C.) formosana, PT, Taiwan, Nantou County, Hueisun Forest Area, slide No.: KA2493f (photo and coll. GR); 95. C. (C.) formosana, PT, Taiwan, Nantou County, Hueisun Forest Area, slide No.: KA043f (coll. HNHM).

FIGURES 114–121. Male 7th and 8th abdominal segments of Cranionycta spp. 114. C. (C.) aurata sp. nov., PT, China, Qinling Mts, N. Ningshan town, slide No.: KA1884m (coll. HS); 115. C. (C.) muscosa sp. nov., HT, China, Taihang Mts, Beijing, Xiaolongmen Nat. Forest Park, slide No.: KA756m (photo and coll. ZISP); 116. C. (C.) oda oda, Russia, Russian Far East, Barabash, slide No.: KA862m (coll. ZMUC); 117. C. (C.) oda transversa, Korea, slide No.: KA1107m (coll. MfN); 118. C. (C.) formosana, HT, Taiwan, Nantou County, Hueisun Forest, MHNG ENTO 60282, slide No.: KA2620m (coll. MHNG); 119. C. (N.) versicolor sp. nov., PT, Nepal, Annapurna Himal, valley of Kali Gandaki, near Tatopani, slide No.: KA2427m (photo and coll. GR); 120. C. (N.) delattini sp. nov., PT, Nepal, Kathmandu valley, Godavari, slide No.: KA1223m (coll. ZSM); 121. C. (N.) warreni sp. nov., PT, Nepal, Kathmandu valley, Godavari, slide No.: KA969m (coll. ZSM). Left side sternite, right side tergite.

FIGURES 130–139. Female 7th abdominal segments of Cranionycta spp. 130. C. (C.) inquieta, China, Prov. Hubei, Dabie Shan, 20 km N of Fengshan, slide No.: KA2527f (coll. PG); 131. C. (C.) muscosa sp. nov., PT, China, Taihang Mts, Beijing, Xiaolongmen Nat. Forest Park, slide No.: KA755f (photo and coll. ZISP); 132. C. (C.) oda oda, Russia, Primorsky Krai, Razdolnoye, slide No.: KA175f (coll. HNHM); 133. C. (C.) oda transversa, South Korea, Prov. North Chungcheong, Mt. Sobaek, Danyang, slide No.: KA1227f (coll. HNHM); 134. C. (C.) formosana, PT, Taiwan, Nantou County, Hueisun Forest Area, slide No.: KA2493f (photo and coll. GR); 135. C. (N.) lobophorus sp. nov., HT, Nepal, Milke Danda, Gursa, slide No.: KA172f (coll. HNHM); 136. C. (N.) versicolor sp. nov., HT, Nepal, Syangja district, Kahule village, slide No.: KA2428f (photo and coll. GR); 137. C. (N.) mailungana sp. nov., HT, Nepal, Ganesh Himal, Mailungkhala, slide No.: KA2529f (coll. PG); 138. C. (N.) delattini sp. nov., HT, Nepal, Kathmandu valley, Godavari, slide No.: KA1224f (coll. GB); 139. C. (N.) warreni sp. nov., HT, Nepal, Kathmandu valley, Godavari, slide No.: KA970f (coll. ZSM). Left side tergite, right side sternite.

FIGURES 21–30. Adults of Cranionycta spp. 21. C. (C.) formosana, HT, male, Taiwan, Nantou County, Hueisun Forest, MHNG ENTO 60282, slide No.: KA2620m (photo and coll. MHNG); 22. C. (C.) formosana, PT, male, Taiwan, Taichung, An-ma-Shan, Hooping, slide No.: KA169m (coll. HNHM); 23. C. (C.) formosana, PT, male, Taiwan, Taichung, An-ma-Shan, Hooping, slide No.: KA170m (coll. HNHM); 24. C. (C.) formosana, PT, female, Taiwan, Nantou County, Hueisun Forest Area, slide No.: KA2493f (coll. GR); 25. C. (N.) lobophorus sp. nov., HT, female, Nepal, Milke Danda, Gursa, slide No.: KA172f (coll. HNHM); 26. C. (N.) mailungana sp. nov., HT, female, Nepal, Ganesh Himal, Mailungkhala, slide No.: KA2529f (coll. PG); 27. C. (N.) versicolor sp. nov., HT, female, Nepal, Syangja district, Kahule village, slide No.: KA2428f (coll. GR); 28. C. (N.) versicolor sp. nov., PT, male, Nepal, Annapurna Himal, valley of Kali Gandaki, near Tatopani, slide No.: KA2427m (coll. GR); 29. C. (N.) delattini sp. nov., HT, female, Nepal, Kathmandu valley, Godavari, slide No.: KA1224f (coll. GB); 30. C. (N.) delattini sp. nov., PT, male, Nepal, Kathmandu valley, Godavari, slide No.: KA1223m (coll. ZSM).

FIGURES 55–60. Male genitalia of Cranionycta spp. 55. C. (C.) aurata sp. nov., HT, China, Prov. Sichuan, near Jiuzhaigou, slide No.: GYP2886 (coll. PG); 56. C. (C.) aurata sp. nov., PT, China, Qinling Mts, N. Ningshan town, slide No.: KA1884m (coll. HS); 57. C. (C.) inquieta, PLT, China, Prov. Chekiang, West Tien-mu-shan (modified after Draudt 1950: pl. 10, fig. 4); 58. C. (C.) inquieta, China, Prov. Jilin, Changbaishan, slide No.: HHL6187 (photo and coll. NEFU); 59. C. (C.) muscosa sp. nov., HT, China, Taihang Mts, Beijing, Xiaolongmen Nat. Forest Park, slide No.: KA756m (photo and coll. ZISP); 60. C. (C.) oda oda, HT, Russia, Primorsky Krai, Suchansky Rudnik [Partizansk] (modified after de Lattin 1949: 111, fig. 3).