Imelda Hewitson, 1870

Zhang, Jing, Cong, Qian, Shen, Jinhui, Opler, Paul A. & Grishin, Nick V., 2021, Genomics-guided refinement of butterfly taxonomy, The Taxonomic Report of the International Lepidoptera Survey 9 (3), pp. 1-55 : 24

Imelda Hewitson, 1870 is a subgenus of Echenais Hübner, [1819]

Despite the differences in appearance, monotypic genus

Echenais Hübner, [1819] (type species Lemonias alphaea

Hübner, [1808], which is Papilio thelephus Cramer, 1775 )

is genetically close to Imelda Hewitson, 1870 (type species

Imelda glaucosmia Hewitson, 1870 , which is a subspecies of Nymphidium mycea Hewitson, 1865 ) (Fig. 23). E.g., COI Fig. 23. Echenais (blue) subgenus Imelda (red).

barcodes of their type species differ by 8.4% (55 bp). Therefore we propose treating Imelda Hewitson, 1870 as a subgenus of Echenais Hübner, [1819] .