Hallonympha maculosa (Bates, 1868), 2021

Zhang, Jing, Cong, Qian, Shen, Jinhui, Opler, Paul A. & Grishin, Nick V., 2021, Genomics-guided refinement of butterfly taxonomy, The Taxonomic Report of the International Lepidoptera Survey 9 (3), pp. 1-55 : 27

Hallonympha maculosa (Bates, 1868) , new combination

Although synonymized with Zabuella Stichel, 1911 (type species Lemonias tenellus Burmeister, 1878 ) by Hall (2018), Hallonympha Penz & DeVries, 2006 (type species Apodemia paucipuncta Spitz, 1930 ) is genetically distant from it (Fig. 28), in agreement with Seraphim et al. (2018). E.g., COI barcodes of the type species of these genera differ by 9.1% (60 bp) and their divergence dates to about 20 Mya ( Seraphim et al. 2018). Therefore, we reinstate Hallonympha as a valid genus. Currently in Callistium (type species Charis cleadas Hewitson, 1866 ), Calydna maculosa Bates, 1868 (type locality Brazil: Amazonas) is not monophyletic with its type species and instead is sister to the type species of Hallonympha (Fig. 28). Their COI barcodes are only 7.4% (49 bp) different. Hence, we propose Hallonympha maculosa (Bates, 1868) comb. nov.