Dalbergia candenatensis (Dennst.) Prain

Adema, F., Ohashi, H. & Sunarno, B., 2016, Notes on Malesian Fabaceae (Leguminosae-Papilionoideae) 17. The genus Dalbergia, Blumea 61 (3), pp. 186-206 : 193

5. Dalbergia candenatensis (Dennst.) Prain — Fig. 1b , 2g

Dalbergia candenatensis (Dennst.) Prain (1901) 49; Merr.(1910) 97; (1923) 294;Backer & Bakh.f.(1964) 614;Verdc.(1979) 295; Sunarno & H.Ohashi (1996) 243; Nyomdham et al. (1997) 45; Sunarno & H.Ohashi (1997) 203; Niyomdham (2002) 130. — Cassia candenatensis Dennst. (1818) 32. — Karin-Tagera, Rheede (1686) 45, t. 25. — Type: Rheede (1686) 45, t. 25.

Dalbergia monosperma Dalzell (1850) 36;Benth.(1852) 256;Miq.(1855) 132; Benth. (1860) 48;Naves & Fern.-Vill.(1880) 67; Perkins (1904) 82. — Type: Dalzell s.n. (K), East India, Malvan Province, Bombay.

Dalbergia torta Graham ex A. Gray (1854) 458, nom. nud.

Dalbergia torta Graham ex Prain (1897a) 120; (1904) 64, t. 42; Ridl. (1922) 591. — Syntypes: Wall . Cat. 5873A (CAL, K), Penang & Singapore; Wall . Cat. 5873B (K), Hb, Finlayson, without locality.

Distribution — India, Ceylon to S China, Cambodia, Vietnam, Thailand, throughout Malesia, N Australia, Solomon Islands, Fiji Islands, New Caledonia.

Habitat & Ecology — Mostly coastal: mangrove, beaches, beach forest, river banks, along roads. Soil: sand, limestone, yellowish soil. Altitude up to 200 m. Flowering: throughout the year; fruiting: February to October, December.

Notes — Dalbergia torta Graham was used by Gray (1854) for a specimen from Fiji, however, without describing the species. Gray gives Wall . Cat. 5789 as the specimen bearing this name. This is, however, according to Prain (1904) an uncor- rected typographical error. Wall . Cat. 5789 is the number given to a set of specimens named Bauhinia spec. None of these specimens is associated with D. torta .

Dalbergia torta was described by Prain (1897) who attributed the name to Graham and gives as specimen Wall . Cat. 5873. This specimen clearly was given the name Dalbergia torta . As Prain (1897) indicates the wrong catalogue number was used by Bentham (1852, 1860), Miquel (1855) and Baker (1879). For D. torta Graham ex Prain Wall. Cat. 5873 is the only candidate as type specimen. However, this Wallich Catalogue number consists of several parts coming from at least two localities. Wall . Cat. 5873 consists of two parts labelled a (A) and b (B). Wall . Cat. 5873A consists of four sheets (3 in K, 1 in CAL) with in total 13 twigs coming from two localities: Penang and Singapore; Wall . Cat. 5873B consists of a single sheet from the herbarium of Finlayson (K) without a locality. For Wall . Cat. 5873 Bentham (1860, sub. D. monosperma ) and Prain (1897, as D. torta ) give Penang as locality, while Ridley (1922, as D. torta ) gives Singapore as locality. It is, however, impossible to tell which twigs were collected in Penang and which in Singapore. We refrain from selecting a lectotype.

Figures

Fig. 1 Wings of Dalbergia species. a. D. junghuhnii Benth., not sculpted; b. D. candenatensis (Dennst.) Prain, sculpted; c. D. pinnata (Lour.) Prain, sculpted (a: SFN 38891 (Sinclair); b: Purseglove & Shah 4633; c: KL 3221). — Drawing by Manon Zuurmond.

Fig. 2 Pods of Dalbergia species. — Type a: a. D. albertisii Prain; b. D. beccarii Prain; c. D. bintuluensis Sunarno & H.Ohashi; d. D. cumingiana Benth.; e. D. kunstleri Prain; f. D. parviflora Roxb. — Type b: g. D. candenatensis (Dennst.) Prain; h. D. menoeides Prain. — Type c: i. D. ferruginea Roxb. (a: NGF 37165 (Womersley); b: Versteeg 1117; c: S 15593 (Ilias Paie); d: BS 18872 (McGregor); e: Hallier 1170; f: SAN 98760 (Amin Sigun); g: Zippelius 70/b; h: Lam 250a; i: SAN102075 (Lee et al.)). — Drawing by Esmée Winkel.