Carbula Stål, 1864

Kaur, R., Singh, D. & Kaur, H., 2013, Taxonomic significance of external genitalia in differentiating four species of genus Carbula Stål (Heteroptera: Pentatomidae) from North India, Journal of Entomology and Zoology Studies 1 (3), No. 3, pp. 33-42 : 41

Key to the studied species of genus Carbula Stål

1 Humeral angles of pronotum prominent, with long acute or subscute spines; scutellum yellowish-green, slightly punctured with black, basal angles with yellowish-green spots and apex impunctate; head somewhat subtriangular; pygophore longer than broad; theca vase-like; paramere with lateral process narrow at base, broad and bifurcated at apex…………………….. 2 - Humeral angles of pronotum subprominent, either with short acute or without spines; scutellum yellowish-brown with dark brown punctation, basal angles with impunctate brownish yellow spots, apex concolorous and puncatae; head somewhat subsqaurish; pygophore as long as broad; theca elongate; paramere with lateral process elongate, subacute at apex without bifurfaction; 3

2. Humeral angles of pronotum with long acute spines; 9 th paratergites elongate, distinctly passing beyond 8 th paratergites; inner margin of blade of paramere slightly convex, obtuse at apex…………………………………. scutellata Distant - Humeral angles of pronotum with obtuse spines; 9 th paratergites broad, just reaching posterior margin of 8 th paratergites; inner margin of blade of paramere strongly convex, subacute at apex…………... socia (Walker)

3. Humeral angles of pronotum with short subacute spines, slightly directed backwards; 9 th paratergites broad, subequal to 8 th paratergites, not passing beyond latter; vesica short, not reaching middle of penial lobes; outer margin of paramere with a distinct medial hump…………………………………………… biguttata (Fabricius) - Humeral angles of pronotum roundish or squarish without spines; 9 th paratergites narrowed at base, lobe-like at apex, passing beyond posterior margin of 8 th paratergites; vesica longer, reaching middle of leaf-like penial lobes; outer margin of paramere without a medial hump…………… insocia (Walker)