Touranella trichosa, Gordana & Ambros, 2018

Golovatch, S. I. & Semenyuk, I. I., 2018, On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda: Polydesmida), Arthropoda Selecta 27 (1), pp. 1-21 : 16-20

publication ID	https://doi.org/10.15298/arthsel.27.1.01
DOI	https://doi.org/10.5281/zenodo.15547203
persistent identifier	https://treatment.plazi.org/id/03D787B3-FFEB-FF84-23BF-F8D9B9AFF9D8
treatment provided by	Felipe
scientific name	Touranella trichosa
status	sp. nov.

Treatment

Touranella trichosa View in CoL sp.n.

Figs 58–73 View Figs 58–61 View Figs 62–64 View Figs 65–68 View Figs 69–71 View Figs 72–73 .

HOLOTYPE ♂, Vietnam, Gia Lai Prov., Kon Ka Kinh National Park , N 14°18′08″, E 108°26′41″, 600–700 m a.s.l., mixed tropical forest, steep slope (up to 45°), stream valley, leaf litter, V.2017, leg. I.I. Semenyuk. GoogleMaps

PARATYPE: 3 ♂♂, 4 ♀♀, same data, together with holotype GoogleMaps .

DIAGNOSIS. Using the latest available key [Golovatch, 2016], this new species keys out to couplet 2 which encompasses several species with densely hirsute collum and following metaterga. However, unlike all of them, T. trichosa sp.n. is totally devoid of a distinct gonofemoral process (at most with a small to vestigial parabasal lobule, j), coupled with a peculiar shape of the solenophore which carries a distinct, acuminate, apical uncus (u) and a couple of characteristic subapical outgrowths (h and n) ( Figs 66–76 View Figs 65–68 View Figs 69–71 View Figs 72–73 View Figs 74–76 ).

NAME. To emphasize the hairy collum and metaterga; adjective.

DESCRIPTION. Both holotype and ♂ paratypes ca 11 mm long, width of midbody pro- and metazonae 0.7 and 0.9 mm, respectively. ♀ paratypes ca 13 mm long, width of midbody pro- and metazonae 1.0–1.1 and 1.3–1.4 mm, respectively. General coloration in alcohol dark brown to brown with venter, legs, tergal hairs and epiproct tip contrasting light greyish or brownish to nearly pallid; tip of antenna pallid ( Figs 58–64 View Figs 58–61 View Figs 62–64 ).

Nearly entire cephalic capsule sparsely setose, setae long, but slightly shorter than tergal hairs ( Figs 59 View Figs 58–61 , 63 View Figs 62–64 ). Antennae rather long and clavate ( Figs 58, 59 View Figs 58–61 , 62, 63 View Figs 62–64 ), in situ extending back behind segment 3 (♂) or 2 (♀) when stretched dorsally; in length, antennomere 3> 2 = 4–6> 1 = 7; interantennal isthmus about as broad as diameter of antennal socket ( Figs 59 View Figs 58–61 , 63 View Figs 62–64 ).

Body clearly moniliform. In width, collum <segments 2–4 <3 <5–17 <head; starting with segment 18, body gradually tapering towards telson ( Figs 59–61 View Figs 58–61 , 63, 64 View Figs 62–64 ). Tegument shining throughout; prozonae very finely shagreened; metaterga and surface below paraterga smooth, only in places very finely striolate longitudinally. Collum and all metaterga beset with long hairs, each hair placed inside a minute pore or on a minute knob ( Figs 58–64 View Figs 58–61 View Figs 62–64 ). Collum broadly and regularly rounded laterally, its paraterga very thin and bordered calluses with a few lateral hairs on minute knobs. Postcollum paraterga very small, set low at about half of body height; paraterga 2 exceptional in being small, declivous, squarish flaps slightly acute-angled and drawn into a nearly sharp tooth anteriorly and into a rounded subrectangular tooth caudally; following paraterga regularly rounded and laterally faintly incised (again due to 3–4 trichomebearing knobs) bulges/calluses typically drawn caudally into a sharp denticle, the latter always far from reaching the posterior margin of metaterga; paraterga slightly thicker in pore-bearing segments than in poreless ones in lateral view ( Figs 58–64 View Figs 58–61 View Figs 62–64 ). Calluses delimited by clear-cut and complete sulci only dorsally, by shorter and less distinct sulci in posterior 1/3–1/2 ventrally. Ozopores small, completely lateral, invisible in dorsal view, lying inside a short longitudinal groove, placed at about 1/3 off caudal corner ( Figs 58 View Figs 58–61 , 62 View Figs 62–64 ). Transverse metatergal sulci complete on segments 5– 17, rather thin, deep, clearly sinuate mid-dorsally, finely beaded at bottom, not reaching the bases of paraterga, but incomplete, abbreviated and traceable as a faint impression on segment 18. Stricture between pro- and metazonae thin and deep, very distinctly ribbed or beaded at bottom down to well below paraterga ( Figs 58, 60–64 View Figs 58–61 View Figs 62–64 ). Axial line evident only in posterior (= postsulcus) halves of postcollum metaterga. Pleurosternal carinae small flaps only in segment 2, thereafter wanting. Epiproct relatively short, subconiform, flattened dorsoventrally, subapical lateral papillae very small, tip faintly concave ( Figs 61 View Figs 58–61 , 64 View Figs 62–64 ). Hypoproct triangular, 1+1 setae strongly separated from each other, borne on minute knobs at caudal margin.

Sterna sparsely setose, cross-impressions faint, axial ones especially so, unmodified except for a high, setose, subtrapeziform lobe between coxae 4 (♂), the lobe supplied with a paramedian pair of remarkably prominent, well separated and caudoventrally directed tubes behind ( Fig. 65 View Figs 65–68 ). Legs long (♂) or short (♀), slender, slightly incrassate and longer in ♂ compared to ♀, ca 1.6–1.7 (♂) or 1.1–1.2 (♀) times as long as midbody height ( Figs 58, 59 View Figs 58–61 , 62, 64 View Figs 62–64 ), devoid of adenostyles, ♂ prefemora not bulged laterally; tarsal brushes missing; in length, femora = tarsi> tibiae> coxae = prefemora = postfemora.

Gonopods ( Figs 59 View Figs 58–61 , 63 View Figs 62–64 , 66–76 View Figs 65–68 View Figs 69–71 View Figs 72–73 View Figs 74–76 ) complex, tripartite as usual; coxite about 1/3 as long as telopodite, subcylindrical, with only a few short setae distolaterally; prefemoral (= densely setose) part of telopodite short, about 1/4 as long as acropodite; femorite rudimenary, immediately distal to prefemoral suclus giving rise to a very long, strong, flat, distally invisible, femoral process (p) with an even longer, flagelliform solenomere (sl) neatly attached to and running along its mesal side; remaining part of solenophore (sph) suberect, longitudinally folded, protecting/sheathing both sl and p nearly all along and leaving only sl tip exposed near a peculiar apical uncus (u) of sph; parabasal part of sph slightly curved dorsad and with a small, sometimes vestigial (broken off?) flap (j); distal 1/3 of sph membranous, u directed mesally, bearing at base a prominent, hyaline, dorsal, apically acuminate lobe (h) and a small, mesal, spatuliform outgrowth (n).