Acrilloscala fortior, Harzhauser & Landau, 2025

Harzhauser, Mathias & Landau, Bernard M., 2025, The Epitoniidae (Gastropoda, Epitonioidea) of the Miocene Central Paratethys Sea-an overwhelming diversity, Zootaxa 5630 (1), pp. 1-142 : 21-22

publication ID

https://doi.org/10.11646/zootaxa.5630.1.1

publication LSID

lsid:zoobank.org:pub:89462472-D932-4255-880F-C6EF15220A31

persistent identifier

https://treatment.plazi.org/id/03D787A4-FFF1-FFF1-FF3F-FC4FFE7B9469

treatment provided by

Plazi

scientific name

Acrilloscala fortior
status

sp. nov.

Acrilloscala fortior sp. nov.

Figs 2I View FIGURE 2 , 8E View FIGURE 8 1 –E View FIGURE 1 2 View FIGURE 2

Type material. Holotype, NHMW 1865 View Materials /0001/1096, SL: 13.4 mm, MD: 4.4 mm, Grund ( Austria), Figs 2I View FIGURE 2 , 8E View FIGURE 8 1 – E View FIGURE 1 2 View FIGURE 2 .

Type locality. Grund ( Austria), North Alpine-Carpathian Foreland Basin .

Type stratum. Grund Formation.

Age. Middle Miocene, early Badenian (Langhian).

Etymology. Fortior (Latin comparative singular of fortis, stronger), referring to the increasing strength of the varices.

Diagnosis. Medium-sized, thin, moderately slender shell with strongly convex whorls, rounded angulation mid-whorl, bearing delicate, close-set lamellae on early teleoconch whorls, becoming wider spaced and more prominent on later whorls, convex base without peribasal cord.

Description. Shell medium-sized, thin, moderately slender (apical angle ~20°). Protoconch unknown. Teleoconch of about 11 whorls, with broad, weakly concave subsutural ramp and rounded angulation mid-whorl. Suture moderately impressed, linear. Sculpture on early teleoconch whorls of delicate, weakly raised, only weakly prosocline close-set lamellae, only partly aligned axially. Lamellae becoming wider spaced and more prominent on later whorls (16: 18: 21). Crests of lamellae only weakly rolled over on last whorl. Tips of lamellae faintly alate, attached to preceding whorl. Few more prominent varices on last whorl formed by amalgamation of several lamellae. Interspaces between lamellae with numerous delicate, barely visible spiral threads. Last whorl attaining 38% of total height, lamellae persisting over base, strongly rolled over, no peribasal cord, base convex delimited by faint basal angulation. Aperture ovate, columella weakly convex. Very short, broad fasciole with distinct growth lines. Outer lip thin, basal lip thin, relatively narrow. Columellar callus forming narrow, adherent rim, parietal callus delicate. Peristome simple. No umbilicus.

Discussion. Acrilloscala terebralis ( Michelin, 1831) (= Acrilloscala degrangei De Boury in Cossmann, 1912), from the Serravallian of France, is much slenderer and has higher whorls (see Ivolas & Peyrot 1900: pl. 2, figs 6–8; Cossmann 1912: pl. 3, figs 17–19; Cossmann & Peyrot 1922: pl. 4, figs 114–115). Acrilloscala sp. 1 , from the Badenian of the Vienna Basin, is also much slenderer and has more prominent lamellae on early teleoconch whorls. Similarly, Acrilloscala geniculata ( Brocchi, 1814) , from the Pliocene of the Mediterranean Sea, differs in its slenderer shape and has distinct, relatively widely spaced lamellae on early whorls (see Bouchet & Warén 1986: figs 1222–1223). Subuliscala subvaricosa ( Cantraine, 1842) as illustrated by Cossmann (1912: pl. 2, figs 16–19) is reminiscent of Acrilloscala fortior sp. nov. in general shape but differs in its more deeply incised suture and the strongly thickened peristome.

Paleoenvironment. At the locality Grund channels formed in middle to outer neritic environments and were filled, with allochthonous assemblages uniting coastal-mudflat faunas with inner neritic ones (Zuschin et al. 2005; Roetzel 2009). The preservation suggests that the species was part of assemblages from middle to outer neritic environments.

Distribution in Central Paratethys. Badenian (Middle Miocene): North Alpine -Carpathian Foreland Basin: Grund ( Austria (hoc opus).

MD

Museum Donaueschingen

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