Papuliscala aquaensis, Harzhauser & Landau, 2025

Harzhauser, Mathias & Landau, Bernard M., 2025, The Epitoniidae (Gastropoda, Epitonioidea) of the Miocene Central Paratethys Sea-an overwhelming diversity, Zootaxa 5630 (1), pp. 1-142 : 10

publication ID

https://doi.org/10.11646/zootaxa.5630.1.1

publication LSID

lsid:zoobank.org:pub:89462472-D932-4255-880F-C6EF15220A31

persistent identifier

https://treatment.plazi.org/id/03D787A4-FFEC-FFED-FF3F-FD43FA2491C9

treatment provided by

Plazi

scientific name

Papuliscala aquaensis
status

sp. nov.

Papuliscala aquaensis sp. nov.

Figs 3A–B View FIGURE 3 , 2A View FIGURE 2 , 4A View FIGURE 4

Type material. Holotype, NHMW 1869 View Materials /0001/0209a, SL: 3.3 mm, MD: 1.1 mm, Baden ( Austria), Figs 3A View FIGURE 3 1 –A View FIGURE 1 2 View FIGURE 2 , 3A View FIGURE 3 , 4A View FIGURE 4 . Paratype, NHMW 1869 View Materials /0001/0209b, SL: 3.2 mm , MD: 0.9 mm, Baden ( Austria), Figs 3B View FIGURE 3 1 –B View FIGURE 1 2 View FIGURE 2 .

Type locality. Baden ( Austria), Vienna Basin .

Type stratum. Baden Formation.

Age. Middle Miocene, middle Badenian (Langhian).

Etymology. Referring to Aquae , the Roman name for the city Baden in Austria,

Diagnosis. Very small, glossy, slender conical with turreted multispiral protoconch and biangulate teleoconch whorls with broad, steep subsutural ramp and prominent axial ribs bearing small nodes at sculptural intersections.

Description. Shell very small, slender conical(apical angle~16–19°). Protoconch slender, turreted of 4.5 smooth, strongly convex whorls, with low nucleus. Last protoconch whorl widening with rounded mid-whorl angulation and delicate, strongly opisthocline axial riblets. Teleoconch of six whorls. Early teleoconch whorls strongly convex with prominent, convex, orthocline axial ribs. Second and third teleoconch whorls successively developing moderately steep, flattened subsutural ramp with angled shoulder, subcylindrical periphery, strongly constricted below basal angulation. Suture deeply impressed, linear. Sculpture of moderately prominent, convex orthocline axial ribs, not aligned axially, separated by wider interspaces (11: 12/13: 12/13). Spiral sculpture of two indistinct cords delimiting angulations forming small nodes at intersections with ribs. Shell surface smooth, glossy. Marked basal angulation delimited by prominent peribasal cord and weaker mid-basal cord. Base nearly flat, smooth. Aperture ovate. Outer lip weakly thickened. Columellar parietal callus delicate, adherent. Peristome simple. No umbilicus.

Discussions. The protoconch with axial riblets on the last whorl confirms placement in Nystiellinae ( Bouchet & Warén 1986: 494). Papuliscala aquaensis sp. nov. is reminiscent of the Pliocene to Recent Mediterranean Papuliscala tavianii Bouchet & Warén, 1986 , which differs in its paucispiral protoconch, more prominent axial ribs and well defined spiral cords (see Bouchet & Warén, 1986: fig. 1163, Landau et al. 2006: pl. 5, figs 3–4). Papuliscala koeneni ( Janssen, 1967) , from the Middle Miocene of the North Sea, also has a multispiral protoconch with axial riblets on the last two whorls but is distinguished from the Paratethyan species by its more prominent spiral cords (see Janssen 1967: pl. 14, fig. 1), with a third cord intercalated between the two primaries on the last two whorls. Papuliscala ambulator Lozouet 1999 , from the late Oligocene of France, has a multispiral protoconch but differs in its higher number of spiral cords and convex whorl profile (see Lozouet 1999: pl. 10, fig. 10).

Paleoenvironment. Occurrences in the Baden Formation of the Vienna Basin suggest middle to outer neritic settings in up to 250 m water depth ( Kranner et al. 2021), which agrees with the environmental requirements of extant species ( Bouchet & Warén 1986; Hoffman et al. 2020).

Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Baden ( Austria) (hoc opus).

MD

Museum Donaueschingen

Kingdom

Animalia

Phylum

Mollusca

Class

Gastropoda

Family

Epitoniidae

Genus

Papuliscala

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