Mazescala kostejana ( Boettger, 1902 )

Harzhauser, Mathias & Landau, Bernard M., 2025, The Epitoniidae (Gastropoda, Epitonioidea) of the Miocene Central Paratethys Sea-an overwhelming diversity, Zootaxa 5630 (1), pp. 1-142 : 91-93

publication ID	https://doi.org/10.11646/zootaxa.5630.1.1
publication LSID	lsid:zoobank.org:pub:89462472-D932-4255-880F-C6EF15220A31
persistent identifier	https://treatment.plazi.org/id/03D787A4-FFBF-FFB8-FF3F-F8B7FAEE9229
treatment provided by	Plazi
scientific name	Mazescala kostejana ( Boettger, 1902 )
status

Treatment

Mazescala kostejana ( Boettger, 1902)

Figs 4G View FIGURE 4 , 33B–C View FIGURE 33 , 36A–C View FIGURE 36

* Scalaria (Hirtoscala) kostejana n. sp. — Boettger 1902: 85.

Scalaria (Hirtoscala) kostejana Boettger — Boettger 1906: 92 .

Hyaloscala kostejensis [sic] Boettger— Cossmann 1912: 170, pl. 5, fig. 3.

Scala (Hyaloscala) kostejana (Boettger) — Zilch 1934: 229, pl. 10, fig. 76.

Scala (Hyaloscala) kostejana (Boettg.) — Báldi 1960: 62, pl. 1, fig. 9.

Scala (Fuscoscala) turtonis alternicostata (Bronn) — Csepreghy-Meznerics 1969a: 22, pl. 4, figs 1, 9 [non Fuscoscala alternicostata ( Bronn, 1831) ].

Scala (Clathrus) kostejana (Boettger, 1901) — Bałuk 1975: 172, pl. 20, figs 15–16.

Epitonium (Hyaloscala) kostejanum (Boettger) — Švagrovský 1981: 140: pl. 42, fig. 5.

Scala (Hyaloscala) kostejana (Boettger, 1901) — Švagrovský 1982: 27: pl. 9, fig. 4.

Type material. Holotype, SMF 380173_12-1446a ( XII 1446 a), specimen illustrated in Zilch (1934: pl. 10, fig. 76), SL: 4.0 mm, MD: 1.2 mm, CoŞteiu de Sus ( Romania), illustrated in Zilch (1934: pl. 10, fig. 80), Fig 36A View FIGURE 36 .

Illustrated material. NHMW 1858/0047/0048, SL: 5.2 mm, MD: 2.5 mm, Porzteich at Břeclav ( Czech Republic), Figs 33B View FIGURE 33 , 36C View FIGURE 36 1 –C View FIGURE 1 2 View FIGURE 2 . NHMW 1859/0019/0108, SL: 3.2 mm, MD: 1.4 mm, Porzteich at Břeclav ( Czech Republic), Figs 4G View FIGURE 4 , 33C View FIGURE 33 , 36B View FIGURE 36 1 –B View FIGURE 1 2 View FIGURE 2 .

Revised description. Shell small, moderately slender (apical angle ~25–28°), faintly coronate. Protoconch turreted of three weakly convex whorls. Teleoconch whorls with weakly concave subsutural ramp and rounded angulation at periphery placed just above mid-whorl. Suture deeply impressed, linear. Early teleoconch whorls with weak, indistinct axial ribs, becoming successively more prominent. Later whorls with numerous, close-set, thin, weakly raised, lamellose to foliaceous, weakly prosocline lamellae, not clearly aligned axially (~18: ~16/20: ~20), crests of lamellae erect. Small, low, vaguely trigonal spines along shoulder. Interspaces glossy with very faint spiral cords, most prominent close to abapical suture and a transition to base. Last whorl with relatively broad, attaining about 45% of total height, with weakly concave subsutural ramp, slightly angled periphery, lamellae persisting over base, strongly rolled over, weak basal angulation, no distinct peribasal cord, moderately narrow, flat to weakly concave base. Aperture subcircular, columella weakly concave. Outer lip thickened by varix. Columellar callus forming narrow rim, detached from base, parietal callus delicate. Umbilicus reduced to narrow chink in subadult specimens; closed in fully grown specimens.

Discussion. Mazescala kostejana ( Boettger, 1902) is characterized by its angled whorls and the ‘tattered’ appearance of the lamellae. It is reminiscent of the recent Australian Mazescala bellicosa ( Hedley, 1907) , which is similar in size (SL: 7.5 mm), sculpture and general outline but differs in its subcylindrical last whorl and the broader base (see Hedley 1907: pl. 67, fig. 18; Iredale 1936: pl. 22, fig. 4). Mazescala thrasys Iredale, 1936 , from Australia, is larger at same growth stage (SL. 15.5 mm), has more teleoconch whorls, is slenderer and has a narrower subsutural ramp on the last whorl (see Iredale 1936: pl. 22, fig. 2). The Australian Mazescala heloris Iredale, 1936 is larger (SL. 13.5 mm) and has a broad subsutural shelf, resulting in a gradate outline (see Iredale 1936: pl. 22, fig. 3). Mazescala koyamai ( Nakayama, 1995) , from Japan, has also somewhat ‘tattered’ lamellae but has a weaker shoulder angulation and Mazescala sakuraii (Kuroda & Habe, 1961) , from Japan, differs in its narrower and shallower subsutural ramp and in its more numerous varices ( Weil et al. 1999: fig. 377). The Japanese Mazescala sawamurai ( Azuma, 1960) has weaker angled whorls and a more convex subsutural ramp ( Weil et al. 1999: fig. 378). The Pliocene to Pleistocene Mediterranean Mazescala salicensis ( Seguenza 1876) is larger (SL: 14.0 mm), has more prosocline lamellae and is slenderer (see Landau et al. 2006: pl. 11, fig. 4).

Paleoenvironment. Probably middle to outer neritic environments.

Distribution in Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Poland) ( Bałuk 1975); Vienna Basin: Porzteich at Břeclav ( Czech Republic) (hoc opus), Borský Mikuláš, Devínska Nová Ves ( Slovakia) ( Švagrovský 1981, 1982); Pannonian Basin: Szokolya ( Hungary) ( Báldi 1960); Bükk Mountains: Balaton ( Hungary) ( Csepreghy-Meznerics 1969a); Făget Basin: CoŞteiu de Sus ( Romania) ( Boettger 1902).

SMF	Forschungsinstitut und Natur-Museum Senckenberg
MD	Museum Donaueschingen
NHMW	Naturhistorisches Museum, Wien

Copyright notice

No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.

Taxonomy

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Family	Epitoniidae
Genus	Mazescala