Hirtoscala steiningeri, Harzhauser & Landau, 2025

Harzhauser, Mathias & Landau, Bernard M., 2025, The Epitoniidae (Gastropoda, Epitonioidea) of the Miocene Central Paratethys Sea-an overwhelming diversity, Zootaxa 5630 (1), pp. 1-142 : 81-82

publication ID	https://doi.org/10.11646/zootaxa.5630.1.1
publication LSID	lsid:zoobank.org:pub:89462472-D932-4255-880F-C6EF15220A31
persistent identifier	https://treatment.plazi.org/id/03D787A4-FFB5-FFB5-FF3F-FAC2FD669469
treatment provided by	Plazi
scientific name	Hirtoscala steiningeri
status	sp. nov.

Treatment

Hirtoscala steiningeri sp. nov.

Figs 27B View FIGURE 27 , 28G View FIGURE 28

Type material. Holotype, HH 42g, Höbarth Museum , Horn ( Austria), SL: 6.2 mm, MD: 3.2 mm, Achberg at Maria Dreieichen ( Austria), Figs 27B View FIGURE 27 1 –B View FIGURE 1 2 View FIGURE 2 , 28G View FIGURE 28 .

Type locality. Achberg at Maria Dreieichen ( Austria), North Alpine Foreland Basin .

Type stratum. Loibersdorf Formation.

Age. Early Miocene, early Eggenburgian (Burdigalian).

Etymology. In honor of Fritz F. Steininger, (Eggenburg, Austria), in recognition of his tremendous impact on Paratethys research.

Diagnosis. Small, solid, moderately slender shell with moderately convex whorls and widely spaced, prominent lamellae with strongly rolled over crests and weak spines along rounded shoulder.

Description. Shell small, solid, moderately slender (apical angle ~20°), weakly coronate. Protoconch and first teleoconch whorls unknown. Teleoconch of moderately convex whorls with rounded shoulder, periphery placed just above mid-whorl. Suture deeply impressed, fenestrate. Sculpture of very prominent, widely spaced, weakly prosocline lamellae, separated by wider interspaces, obliquely aligned axially (?: 8: 8), forming short, indistinct spines along shoulder. Crests of lamellae fragile, strongly rolled over. Interspaces between lamellae smooth. Last whorl attaining about 40% of total height, without basal angulation or peribasal cord, lamellae strongly rolled over on base. Fasciole very short, broad, with prominent growth lines. Aperture ovate, columella weakly concave. Columellar callus and parietal callus continuous, columellar callus forming narrow rim, attached to fasciole. Peristome thickened, duplex. No umbilicus.

Discussion. Hirtoscala steiningeri sp. nov. co-occurs with Hirtoscala ceruchus sp. nov. and Hirtoscala uncinata sp. nov., which all fall in the same size class. Hirtoscala uncinata differs clearly in its strongly coronate shoulder whereas H. ceruchus differs in its uniformly broad lamellae, which lack spines and the low position of the periphery.

The Pliocene Mediterranean Hirtoscala spinifera ( Seguenza, 1876) might be a closely related species but differs in its much larger size and broader shell (see Landau et al. 2006: pl. 9, figs 11–16, Chirli 2009: pl. 14, figs 1–7).

Distribution in Central Paratethys. Eggenburgian (Early Miocene): North Alpine Foreland Basin: Achberg at Maria Dreieichen ( Austria) ( Steininger et al. 1971).