Hirtoscala uncinata, Harzhauser & Landau, 2025

Harzhauser, Mathias & Landau, Bernard M., 2025, The Epitoniidae (Gastropoda, Epitonioidea) of the Miocene Central Paratethys Sea-an overwhelming diversity, Zootaxa 5630 (1), pp. 1-142 : 82-83

publication ID	https://doi.org/10.11646/zootaxa.5630.1.1
publication LSID	lsid:zoobank.org:pub:89462472-D932-4255-880F-C6EF15220A31
persistent identifier	https://treatment.plazi.org/id/03D787A4-FFB4-FFB6-FF3F-FDD3FD6696A5
treatment provided by	Plazi
scientific name	Hirtoscala uncinata
status	sp. nov.

Treatment

Hirtoscala uncinata sp. nov.

Figs 28H View FIGURE 28 , 31A–F View FIGURE 31

Scala (Spiniscala) merignacensis De Boury, 1912 — Steininger et al. 1971: 385, pl. 3, figs 7–8 [non Hirtoscala merignacensis (de Boury in Cossmann, 1912)].

Type material. Holotype, HH42, Höbarth Museum , Horn ( Austria), SL: 6.9 mm, MD: 3.4 mm, Achberg at Maria Dreieichen ( Austria), Figs 28H View FIGURE 28 , 31D View FIGURE 31 1 –D View FIGURE 1 2 View FIGURE 2 . Paratypes, HH42a, Höbarth Museum , Horn ( Austria), SL: 8.0 mm , MD: 4.6 mm, Achberg at Maria Dreieichen ( Austria), Figs 31A View FIGURE 31 1 –A View FIGURE 1 2 View FIGURE 2 . HH42b, Höbarth Museum, Horn ( Austria), SL: 8.1 mm , MD: 3.2 mm, Achberg at Maria Dreieichen ( Austria), Figs 31B View FIGURE 31 1 –B View FIGURE 1 2 View FIGURE 2 . HH42c, Höbarth Museum, Horn ( Austria), SL: 8.0 mm , MD: 3.8 mm, Achberg at Maria Dreieichen ( Austria), Figs 31C View FIGURE 31 1 –C View FIGURE 1 2 View FIGURE 2 . HH42d, Höbarth Museum, Horn ( Austria), SL: 7.3 mm , MD: 3.5 mm, Achberg at Maria Dreieichen ( Austria), Fig. 31E View FIGURE 31 . HH42e, Höbarth Museum, Horn ( Austria), SL: 8.6 mm , MD: 4.0 mm, Achberg at Maria Dreieichen ( Austria), Fig. 31F View FIGURE 31 .

Type locality. Achberg at Maria Dreieichen ( Austria), North Alpine Foreland Basin .

Type stratum. Loibersdorf Formation.

Age. Early Miocene, early Eggenburgian (Burdigalian).

Etymology. Uncinatus (Latin for hook-like), referring to the axial sculpture.

Diagnosis. Small, solid, moderately slender, gradate and strongly coronate shell with very prominent, broad, strongly rolled over lamellae forming long, solid spines pointed in adapical direction.

Description. Shell small, solid, moderately slender (apical angle ~20–25°), gradate, strongly coronate. Protoconch and first teleoconch whorls unknown. Teleoconch of weakly convex, subcylindrical whorls with broad, concave subsutural shelf and angled shoulder, with periphery placed mid-whorl. Suture deeply impressed, largely obscured by lamellae. Sculpture of very prominent, moderately close-set, only weakly prosocline lamellae, separated by slightly wider interspaces, not aligned axially (?: ~10: ~11/14), forming very prominent, adapically pointed spines along shoulder. Crests of lamellae strongly rolled over. Interspaces between lamellae smooth. Last whorl attaining about 45% of total height, without basal angulation nor peribasal cord, lamellae strongly rolled over on base. Fasciole very short, broad, with prominent growth lines. Aperture ovate, columella weakly concave. Columellar and parietal callus continuous, columellar callus forming narrow rim, attached to fasciole. Peristome thickened, duplex. No umbilicus.

Discussion. Hirtoscala uncinata sp. nov. is characterized by its extraordinarily prominent lamellae with long, solid spines. This species was confused by Steininger et al. (1973) with the coeval Hirtoscala merignacensis (de Boury in Cossmann, 1912) from the Burdigalian of the Aquitaine Basin in France, which differs in its broader conical outline and fewer lamellae (see Cossmann 1912: pl. 1, figs 30–31; Cossmann & Peyrot 1922: pl. 3, figs 39–42). Hirtoscala benoisti de Boury, 1912 , from the Burdigalian of the Aquitaine Basin, has even fewer lamellae than H. merignacensis and differs from the Paratethyan species in its slightly inflated last whorl ( Cossmann 1912: pl. 1, figs 33–34; Cossmann & Peyrot 1922: pl. 3, figs 69–70). Hirtoscala hastata sp. nov. has more numerous lamellae and less prominent spines.

The extant Hirtoscala venosum (G.B. Sowerby II, 1844) , from the western Atlantic is comparable in its strongly coronate lamellae and gradate spire but differs in its broader shell ( Weil et al. 1999: fig. 28).

Paleoenvironment. Shallow marine, inner neritic.

Distribution in Central Paratethys. Eggenburgian (Early Miocene): North Alpine Foreland Basin: Achberg at

Maria Dreieichen ( Austria) ( Steininger et al. 1971).