Hyaloscala rainesi, Harzhauser & Landau, 2025

Harzhauser, Mathias & Landau, Bernard M., 2025, The Epitoniidae (Gastropoda, Epitonioidea) of the Miocene Central Paratethys Sea-an overwhelming diversity, Zootaxa 5630 (1), pp. 1-142 : 85-86

publication ID

https://doi.org/10.11646/zootaxa.5630.1.1

publication LSID

lsid:zoobank.org:pub:89462472-D932-4255-880F-C6EF15220A31

persistent identifier

https://treatment.plazi.org/id/03D787A4-FFB1-FFB1-FF3F-FA59FF60959D

treatment provided by

Plazi

scientific name

Hyaloscala rainesi
status

sp. nov.

Hyaloscala rainesi sp. nov.

Figs 28J View FIGURE 28 , 32B View FIGURE 32 1 –B View FIGURE 1 2 View FIGURE 2

Type material. Holotype, NHMW 2024 View Materials /0278/0006, SL: 5.2 mm, MD: 2.3 mm, Bad Vöslau ( Austria), Figs 28J View FIGURE 28 , 32B View FIGURE 32 1 –B View FIGURE 1 2 View FIGURE 2 .

Type locality. Bad Vöslau ( Austria), Vienna Basin .

Type stratum. Baden Formation.

Age. Middle Miocene, middle Badenian (Langhian).

Etymology. In honor of Bret Raines (Victorville, California, USA), in recognition of his contributions in Epitoniidae systematics.

Diagnosis. Small, moderately broad, stocky conical, weakly gradate, faintly coronate, with narrow subsutural ramp, weak shoulder angulation, moderately close-set, moderately raised lamellae, obliquely aligned axially, sub-angled base, subcircular aperture.

Description. Shell small, translucent, moderately broad, stocky conical (apical angle ~32°), weakly gradate, faintly coronate. Protoconch and early teleoconch whorls unknown. Teleoconch of at least seven whorls. Early whorls with slight mid-whorl angulation, later whorls with narrow, weakly concave subsutural ramp and convex periphery placed mid-whorl. Suture deeply impressed. Sculpture of moderately close-set, moderately raised, only weakly prosocline lamellae, obliquely aligned axially (12: 16: 16), forming low, trigonal spines along shoulder, fused across suture. Crests of lamellae only weakly rolled over. Interspaces between lamellae smooth, glossy. Last whorl attaining 45% of total height, faintly coronate, lamellae persisting over base, strongly rolled over, base sub-angled, no peribasal cord, flattened. Aperture subcircular, columella weakly concave. Very short, broad fasciole with distinct growth lines. Columellar callus forming narrow rim, attached to fasciole, parietal callus delicate. Peristome simple, slightly flaring but not thickened. No umbilicus.

Discussion. The extant Hyaloscala opposita (de Boury, 1921) , from South Africa, is exceptionally similar and differs only in its slightly weaker shoulder angulation and more convex base. (for holotype see https://science.mnhn. fr/institution/mnhn/collection/im/item/2000-4366). This African species was placed in Sodaliscala by de Boury (1921), but Sodaliscala has spiral cords in the interspaces between the lamellae ( Kilburn 1985: 312). Consequently, the species was transferred to Hyaloscala by Kilburn (1985: 293) and Weil et al. (1999: 54). The Atlanto-Mediterranean Hyaloscala clathratula Kanmacher, 1798 has a much weaker shoulder angulation (see MolluscaBase eds 2924i). Hyaloscala miotrinacria ( Sacco, 1891) , from the Late Miocene of Tetti Borelli ( Italy), differs in its greater number of lamellae and broader early teleoconch whorls (see Sacco 1891: pl. 1, fig. 42; Ferrero Mortara et al. 1984: pl. 3, fig. 13). Hyaloscala micans Cossmann & Peyrot, 1922 , from the Burdigalian of France, is comparably stout with broader spire whorls and larger last whorl (see Cossmann & Peyrot 1922: pl. 4, figs 86–87). Hyaloscala exilis de Boury, 1912 , from the Burdigalian of France, ( Cossmann & Peyrot 1922: pl. 4, fig, 25), is much slenderer and has more numerous, delicate lamellae.

Paleoenvironment. Occurrences in the Baden Formation of the Vienna Basin suggest middle to outer neritic settings in up to 250 m water depth ( Kranner et al. 2021).

Distribution in Central Paratethys. Badenian (Middle Miocene): Vienna Basin: Bad Vöslau ( Austria) (hoc opus).

MD

Museum Donaueschingen

Kingdom

Animalia

Phylum

Mollusca

Class

Gastropoda

Family

Epitoniidae

Genus

Hyaloscala

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