Hirtoscala diplommatinaformis, Harzhauser & Landau, 2025
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publication ID |
https://doi.org/10.11646/zootaxa.5630.1.1 |
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publication LSID |
lsid:zoobank.org:pub:89462472-D932-4255-880F-C6EF15220A31 |
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persistent identifier |
https://treatment.plazi.org/id/03D787A4-FFAE-FFA8-FF3F-FED7FE379481 |
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treatment provided by |
Plazi |
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scientific name |
Hirtoscala diplommatinaformis |
| status |
sp. nov. |
Hirtoscala diplommatinaformis sp. nov.
29B 1 –B 2
Type material. Holotype, NHM 1862/0029/0081, SL: 8.5 mm, MD: 5.6 mm, Rudice ( Czech Republic), Figs 29B View FIGURE 29 1 – B View FIGURE 1 2 View FIGURE 2 .
Type locality. Rudice ( Czech Republic), Carpathian Foreland Basin .
Type stratum. unknown.
Age. Middle Miocene, middle Badenian (Langhian).
Etymology. Referring to its superficial similarity with some species of the terrestrial gastropod Diplommatina .
Diagnosis. Small, very broad, stout conical, gradate, coronate shell with prominent shoulder, close-set, strongly raised, thin lamellae, partly obliquely aligned axially, distinct chink between fasciole and columellar callus.
Description. Shell small, very broad, stout conical, gradate, coronate (apical angle ~41°). Protoconch and early teleoconch whorls unknown. Teleoconch of relatively low whorls, with shallow, weakly convex subsutural ramp, prominent shoulder, periphery placed above mid-whorl. Suture distinctly impressed, weakly undulating. Sculpture of close-set, strongly raised, lamellose, weakly prosocline lamellae, partly obliquely aligned axially (?: 17: 16). Lamellae forming short, trigonal spines along shoulder. Second row of weaker spines at suture, separated from shoulder spines by concavity. Interspaces between lamellae glossy, smooth. Last whorl attaining about 45% of total height, lamellae persisting over base with strongly rolled over crests, resulting in flattish appearance, distinct basal angulation coinciding with slightly angled lamellae, no peribasal cord, base weakly concave. Short, broad fasciole with prominent growth lines. Aperture subcircular, columella concave. Outer lip thickened by varix, basal lip broad. Columellar callus forming narrow rim separated from fasciole by deep chink; parietal callus delicate. Peristome duplex. No umbilicus.
Discussion. We place this peculiar species in Hirtoscala based on its slightly coronate lamellae and smooth interspaces. Hirtoscala diplommatinaformis sp. nov. is reminiscent of a squat morph of Hirtoscala muricatoides ( Sacco, 1891) , from the Mediterranean Pliocene, and Hirtoscala pantanellii (de Boury in Cossmann, 1912), from the Mediterranean Pleistocene (see Cossmann 1912: pl. 2, fig. 26; Ferrero Mortara et al. 1984: pl. 5, fig. 3; Landau et al. pl. 8, figs 1–2, pl. 9, figs 1–6). However, all three species are somewhat untypical for Hirtoscala due to their weak basal disc. Hirtoscala muricatoides ( Sacco, 1891) and Hirtoscala pantanellii (de Boury in Cossmann, 1912) were placed in Gyroscala by Cossmann (1912: 48) but lack the broad peribasal cord and the axially fused lamellae typical of that genus.
Hirtoscala diplommatinaformis sp. nov. is superficially reminiscent of Tioria Maxwell, 1928 [type species Cirsotrema (Tioria) youngi Marwick, 1928 ; Paleocene/Eocene, New Zealand]. Tioria is characterized by its gradate shell with broad subsutural ramp and strongly rolled over crests of the lamellae on the base and a chink between fasciole and columellar callus; it lacks a peribasal cord. However, it differs from Hirtoscala diplommatinaformis sp. nov. in its spiral sculpture and the detached parietal lip. Tioria was described so far only from the Paleogene of New Zealand and was documented only by its type species [see Beu & Maxwell (1990: 96) and Beu & Raine (2009)].
Paleoenvironment. Probably middle to outer neritic based on co-occurring assemblage. (own data M.H.).
Distribution in Central Paratethys. Badenian (Middle Miocene): Carpathian Foreland Basin: Rudice ( Czech Republic) (hoc opus).
| MD |
Museum Donaueschingen |
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