Hirtoscala miofrondiculoides ( Sacco, 1891 )

Harzhauser, Mathias & Landau, Bernard M., 2025, The Epitoniidae (Gastropoda, Epitonioidea) of the Miocene Central Paratethys Sea-an overwhelming diversity, Zootaxa 5630 (1), pp. 1-142 : 79-80

publication ID	https://doi.org/10.11646/zootaxa.5630.1.1
publication LSID	lsid:zoobank.org:pub:89462472-D932-4255-880F-C6EF15220A31
persistent identifier	https://treatment.plazi.org/id/03D787A4-FFAB-FFAB-FF3F-F924FEB890CA
treatment provided by	Plazi
scientific name	Hirtoscala miofrondiculoides ( Sacco, 1891 )
status

Treatment

Hirtoscala miofrondiculoides ( Sacco, 1891)

Figs 28E View FIGURE 28 , 30D–F View FIGURE 30

Scalaria pseudoscalaria Brocc.— Hörnes 1848: 22 [non Hirtoscala pseudoscalaris ( Brocchi, 1814) ].

Scalaria clathratula Turt.—Hörnes 1856: 475 View in CoL (pars), pl. 46, figs 8a–d [non Hyaloscala clathratula ( Kanmacher, 1798) View in CoL , nec Turton, 1806).

[ Scalaria View in CoL ] H [irtoscala]. miofrondiculoides — Sacco 1891: 26 [nov. nom. pro Scalaria clathratula Hörnes, 1856 View in CoL , non Kanmacher, 1798, nec Turton, 1806].

Clathrus parilis De Boury nom. mut.— Cossmann 1912: 172, pl. 5, fig. 5.

Scala (Spiniscala) frondicula Wood var. spinosa Bon.— Meznerics 1933: 337 View in CoL , pl. 14, figs 5a–b [non Hirtoscala spinosa ( Sacco, 1891) ].

S [cala]. (S [cala].) frondicula View in CoL spinosa Bon.— Sieber 1958: 129 [non Hirtoscala spinosa ( Sacco, 1891) ].

Scala (Clathrus) spinosa Bonelli, 1826 View in CoL — Bałuk 1975: 171, pl. 20, figs 17–18 [non Hirtoscala spinosa ( Sacco, 1891) ]. Epitonium miofrondiculoides ( Sacco, 1891) — Mandic & Harzhauser 2003: 97, pl. 1, figs 2–3.

Epitonium miofrondiculoides ( Sacco, 1891) — Landau et al. 2013: 138, pl. 20, fig. 8.

non Scalaria clathrata View in CoL — Eichwald 1852: pl. 10, fig. 20 [non Kanmacher, 1798, nec Turton, 1806] [= Hirtoscala sp. ].

non Scala spinosa Bonelli— Friedberg 1938: 83 View in CoL , text-fig. 20 [= Hirtoscala hastata sp. nov.].

non Scala (Hirtoscala) frondicula spinosa Bon.— Strausz 1954: 19 View in CoL , pl. 1, fig. 9 [= Hirtoscala holcovae sp. nov.].

Type material. Holotype, NHMW 2024 View Materials /0282/0001, SL: 9.3 mm, MD: 3.7 mm, Steinebrunn ( Austria), illustrated in Hörnes (1856: pl. 46, figs 8a–d), Figs 28E View FIGURE 28 , 30D View FIGURE 30 1 –D View FIGURE 1 2 View FIGURE 2 .

Illustrated material. NHMW 2024 View Materials /0278/0004, SL: 9.3 mm , MD: 4.1 mm, Bad Vöslau ( Austria), Figs 30E View FIGURE 30 1 – E View FIGURE 1 2 View FIGURE 2 . NHMW 2024 View Materials /0278/0005, SL: 4.7 mm , MD: 2.2 mm, Bad Vöslau ( Austria), Figs, 30F.

Revised description. Shell small, moderately slender (apical angle 24°), faintly coronate. Protoconch and early teleoconch whorls unknown. Teleoconch of at least eight convex whorls, with periphery placed below mid-whorl. Narrow, weakly concave subsutural shelf. Suture moderately impressed. Sculpture of moderately closely set, robust, strongly raised, weakly prosocline lamellae, obliquely aligned axially (11: 11: 11), adapical tips slightly pointed close to suture. Crests of lamellae strongly rolled over. Interspaces between lamellae smooth. Last whorl attaining about 40% of total height, lamellae persisting over base, strongly rolled over, no peribasal cord nor basal angulation, base slightly convex. Fasciole very short, broad, with prominent growth lines. Aperture ovate, columella weakly concave. Outer lip thickened by varix, basal lip broad. Columellar callus forming moderately broad rim, slightly detached from base, parietal callus delicate. Peristome duplex but inner layer covers outer layer largely. No umbilicus.

Synonyms. Clathrus parilis De Boury in Cossmann, 1912, was introduced by Cossmann (1912) from CoŞteiu de Sus ( Romania). The holotype, illustrated in Cossmann (1912: pl. 5, fig. 5) does not differ from Hirtoscala miofrondiculoides ( Sacco, 1891) . No good illustration of this species, however, was available to Cossmann (1912) at that time.

Discussion. The holotype was mixed by Hörnes (1856) with specimens described herein as Parviscala lacinia sp. nov. Whilst the latter is quite frequent at Steinebrunn ( Austria) with more than 50 specimens, Hirtoscala miofrondiculoides is very rare. They are clearly distinguished by the gradate spire and weak spiral cords of P. lacinia . Hirtoscala hastata sp. nov. differs from H. miofrondiculoides in its subcylindrical whorls and slightly wider subsutural shelf. The Pliocene Mediterranean Hirtoscala proxima (de Boury, 1890) might be a closely related species but is slenderer and has a slightly deeper incised suture (see de Boury 1890, pl. 4, fig. 9; Landau et al. 2006: pl. 11, figs 1–2). Scalaria clathrata sensu Eichwald (1852 : pl. 10, fig. 20) differs in its strongly convex whorls and its varices are not fused across the suture. The status of this species from Ukraine remains unclear.

Scalaria spinosa Bonelli refers to a name used in an unpublished catalogue for the collection in Torino ( Italy) written by Franco Andrea Bonelli (1784–1830) around the year 1826. The name was made available by Sacco (1891: 26, pl. 1, fig. 38), who illustrated a specimen from the Tortonian of S. Agata ( Italy). Hirtoscala spinosa ( Sacco, 1891) differs from H. miofrondiculoides in its slightly gradate spire and subcylindrical whorls.

Paleoenvironment. The occurrence at Steinebrunn ( Austria) suggests shallow marine environments with sea grass (own data M.H.).

Distribution in Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Korytnica ( Poland) ( Bałuk 1975); Vienna Basin: Baden, Bad Vöslau, Steinebrunn ( Austria) (Hörnes 1856); Făget Basin: CoŞteiu de Sus ( Romania) ( Cossmann 1912).

Proto-Mediterranean Sea. Serravallian (Middle Miocene): Karaman Basin: Seyithasan ( Turkey) ( Landau et al. 2013).

MD	Museum Donaueschingen

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Taxonomy

Kingdom	Animalia
Phylum	Mollusca
Class	Gastropoda
Family	Epitoniidae
Genus	Hirtoscala