Hirtoscala hastata, Harzhauser & Landau, 2025

Harzhauser, Mathias & Landau, Bernard M., 2025, The Epitoniidae (Gastropoda, Epitonioidea) of the Miocene Central Paratethys Sea-an overwhelming diversity, Zootaxa 5630 (1), pp. 1-142 : 77-78

publication ID	https://doi.org/10.11646/zootaxa.5630.1.1
publication LSID	lsid:zoobank.org:pub:89462472-D932-4255-880F-C6EF15220A31
persistent identifier	https://treatment.plazi.org/id/03D787A4-FFA9-FFA9-FF3F-FC8CFB0E9144
treatment provided by	Plazi
scientific name	Hirtoscala hastata
status	sp. nov.

Treatment

Hirtoscala hastata sp. nov.

Figs 28C View FIGURE 28 , 30A–B View FIGURE 30

Scala spinosa Bonelli— Friedberg 1938: 83 View in CoL , text-fig. 20 [non Hirtoscala spinosa ( Sacco, 1891) ].

Illustrated material. Holotype, NHMW 2022 View Materials /0290/0169, Vienna / Ottakring ( Austria), SL: 6.2 mm, MD: 2.3 mm, Figs 30B View FIGURE 30 1 –B View FIGURE 1 2 View FIGURE 2 . Paratype, NHMW 2024 View Materials /0278/0003, SL: 7.4 mm , MD: 2.8 mm, Bad Vöslau ( Austria), Figs 28C View FIGURE 28 , 30A View FIGURE 30 1 –A View FIGURE 1 2 View FIGURE 2 .

Type locality. Vienna / Ottakring ( Austria), Vienna Basin .

Type stratum. Baden Formation.

Age. Middle Miocene, Badenian (Langhian).

Etymology. Hastatus (Latin for armed with a spear), referring to the slender outline.

Diagnosis Small, moderately slender, turriculate, coronate shell with narrow subsutural shelf, moderately close-set, robust, strongly raised, weakly prosocline lamellae with low, trigonal spines at shoulder, smooth interspaces.

Description. Shell small, moderately slender, turriculate (apical angle ~20–22°), coronate. Protoconch and early teleoconch whorls unknown. Teleoconch of at least eight whorls, with narrow subsutural shelf and weakly convex to subcylindrical periphery. Suture deeply impressed. Sculpture of moderately close-set, robust, strongly raised, weakly prosocline lamellae, aligned axially (13: 13: 14), adapical tips forming low, trigonal spines along shoulder. Crests of lamellae slightly rolled over. Interspaces between lamellae smooth, glossy. Last whorl attaining about 41% of total height, lamellae persisting over base, becoming strongly rolled over, flat and broad, no peribasal cord nor basal angulation but lamellae faintly angled at transition into flattish base. Very short, broad fasciole with prominent growth lines. Aperture subcircular, columella weakly concave. Outer lip thickened by prominent varix. Columellar callus forming broad rim, detached from base, parietal callus delicate. Peristome duplex but inner layer covering outer layer largely. No umbilicus.

Discussion. Hirtoscala miofrondiculoides ( Sacco, 1891) differs in its fewer and broader varices, which are distinctly fused over the suture. The extant eastern Atlantic Spiniscala trevelyana ( Johnston, 1841) is reminiscent of the Paratethyan species but differs in its wider shell and has an open umbilicus ( Weil et al. 1999: fig. 92). ‘ Epitonium trevelyanum’ from the Serravallian of the Karaman Basin ( Turkey), is also a Hirtoscala but lacks a coronate shoulder (see Landau et al. pl. 20, fig. 10). Hirtoscala spinosa ( Sacco, 1891) , from the Tortonian of S. Agata ( Italy) might be closely related but differs in in having fewer lamellae (see Sacco 1891: pl. 1, fig 38).

Hirtoscala hastata sp. nov. is exceptionally similar to the extant Hirtoscala tenebrosa (G.B. Sowerby II, 1903) from South Africa, which differs mainly in its slightly broader and more convex whorls and deeper suture (see Kilburn 1985: figs 94–95; Weil et al. 1999: fig. 142). The Pliocene Mediterranean Hirtoscala spinifera ( Seguenza, 1876) differs in its fewer, much broader lamellae (see Landau et al. 2006: pl. 9, figs 11–16, Chirli 2009: pl. 14, figs 1–7).

Parviscala lacinia sp. nov. and Parviscala paratethyca sp. nov. are superficially similar but differ in the presence of spiral sculpture in the axial interspaces and have fewer lamellae. Moreover, H. hastata has a narrower subsutural ramp with weaker spines compared to Parviscala lacinia sp. nov.

Paleoenvironment. Occurrences in the Baden Formation of the Vienna Basin suggest middle to outer neritic settings in up to 250 m water depth ( Kranner et al. 2021).

Distribution in Central Paratethys. Badenian (Middle Miocene): Korytnica Basin: Jawor at Korytnica ( Poland) ( Friedberg 1938); Vienna Basin: Bad Vöslau, Vienna /Ottakring ( Austria) (hoc opus).