Parviscala lacinia, Harzhauser & Landau, 2025

Harzhauser, Mathias & Landau, Bernard M., 2025, The Epitoniidae (Gastropoda, Epitonioidea) of the Miocene Central Paratethys Sea-an overwhelming diversity, Zootaxa 5630 (1), pp. 1-142 : 108-110

publication ID	https://doi.org/10.11646/zootaxa.5630.1.1
publication LSID	lsid:zoobank.org:pub:89462472-D932-4255-880F-C6EF15220A31
persistent identifier	https://treatment.plazi.org/id/03D787A4-FF8E-FF89-FF3F-FB3EFADE9309
treatment provided by	Plazi
scientific name	Parviscala lacinia
status	sp. nov.

Treatment

Parviscala lacinia sp. nov.

Figs 4N–O View FIGURE 4 , 40B View FIGURE 40 , 41A–H View FIGURE 41

Scalaria clathratula Turt.—Hörnes 1856: 475 View in CoL (pars) [non Hyaloscala clathratula ( Kanmacher, 1798) View in CoL , nec Turton, 1806).

Spiniscala spinosa (Bonelli) — Cossmann 1912: 169, pl. 5, figs 2 [non Hirtoscala spinosa ( Sacco, 1891) ].

Scala Duboisi Friedb.— Friedberg 1928: 605 View in CoL , pl. 38, fig. 22 [non ‘Scala’ duboisi Friedberg, 1914 .].

Scala duboisi Friedberg, 1914 — Zelinskaya et al. 1968: 173, pl. 41, fig. 11 [non ‘Scala’ duboisi Friedberg, 1914 ].

Scala (Clathrus) frondicula spinosa Bonelli, 1826 View in CoL — Strausz 1966: 175, pl. 1, fig. 34 [non Hirtoscala spinosa ( Sacco, 1891) ].

Scala (Fusoscala) frondicula (Wood) View in CoL — Atanacković 1969: 200, pl. 9, fig. 14 [non Hirtoscala frondicula ( Wood, 1848) ].

Scala (Spiniscala) frondicuala spinosa ( Bonelli, 1826) View in CoL — Atanacković 1985: 115, pl. 27, figs 15–16 [non Hirtoscala spinosa ( Sacco, 1891) ].

Type material. Holotype, NHMW 2018 View Materials /0248/0551a, SL: 4.9 mm, MD: 2.0 mm, CoŞteiu de Sus ( Romania), Figs 41A View FIGURE 41 1 –A View FIGURE 1 2 View FIGURE 2 . Paratypes, NHMW 2018 View Materials /0248/0551b, SL: 3, 1 mm , MD: 1.7 mm, CoŞteiu de Sus ( Romania) , Figs 4N View FIGURE 4 , 41C View FIGURE 41 1 –C View FIGURE 1 2 View FIGURE 2 . NHMW 2018 View Materials /0248/0551c, SL: 5.2 mm , MD: 2.1 mm, CoŞteiu de Sus ( Romania) , Figs 41D View FIGURE 41 1 –D View FIGURE 1 2 View FIGURE 2 . NHMW 1855 View Materials /0045/0206a, SL: 6.1 mm , MD: 2.6 mm, Steinebrunn ( Austria) , Figs 39B View FIGURE 39 , 41E View FIGURE 41 1 –E View FIGURE 1 2 View FIGURE 2 . NHMW 1855 View Materials /0045/0206b, SL: 6.8 mm , MD: 2.9 mm, Steinebrunn ( Austria) , Figs 40B View FIGURE 40 1 –B View FIGURE 1 2 View FIGURE 2 . NHMW 1887 View Materials /0018/0017, SL: 7.8 mm , MD: 4.1 mm, Steinebrunn ( Austria) . NHMW 2024 View Materials /0278/0016, SL: 4.4 mm , MD: 1.7 mm, Bad Vöslau ( Austria) , Figs 41F View FIGURE 41 1 – F View FIGURE 1 2 View FIGURE 2 . NHMW 2024 View Materials /0278/0017, SL: 3.1 mm , MD: 1.4 mm, Bad Vöslau ( Austria) , Fig. 41G View FIGURE 41 . NHMW 2024 View Materials /0278/0018, SL: 3.7 mm , MD: 1.6 mm, Bad Vöslau ( Austria) , Figs 41H View FIGURE 41 1 –H View FIGURE 1 2 View FIGURE 2 . NHMW 1887 View Materials /0018/0017, SL: 7.8 mm , MD: 4.1 mm, Steinabrunn ( Austria) , Fig. 4O View FIGURE 4 .

Additional material. 52 spec., NHMW 1864 View Materials /0001/0694, Steinebrunn ( Austria) .

Type locality. CoŞteiu de Sus ( Romania), Făget Basin .

Type stratum. Silt and clay of the Dej Formation.

Age. Middle Miocene, Badenian (Langhian).

Etymology. After lacinia (Latin for corner, end, tip, point), referring to the small shoulder spines; noun in apposition.

Diagnosis. Small, moderately broad, turriculate, gradate, coronate with convex to subcylindrical whorls, narrow subsutural shelf, robust, strongly raised, weakly prosocline lamellae forming short spines along shoulder, delicate spiral sculpture.

Description. Shell small, moderately broad, gradate (apical angle ~27–29°), coronate. Protoconch turreted of 4.5 smooth, weakly convex whorls, last whorl with marked mid-whorl angulation, nucleus moderately elevated dome. Teleoconch of five whorls, with narrow subsutural shelf and weakly convex to subcylindrical periphery. Suture deeply impressed. Sculpture of moderately close-set, robust, strongly raised, weakly prosocline lamellae, only partly obliquely aligned axially (9/11: 13: 13), adapical tips forming robust, trigonal, strongly pointed spines along shoulder. Crests of lamellae slightly rolled over on spire whorls, strongly rolled over on last whorl. Interspaces between lamellae with broad, low spiral cords. Last whorl attaining about 40% of total height, lamellae persisting over base, strongly rolled over, flattish, no peribasal cord, base slightly concave. Aperture ovate to subcircular, columella weakly concave. Outer lip thickened by prominent varix. Columellar callus forming broad rim, detached from base, parietal callus delicate. Peristome duplex with inner layer covering outer layer largely. No umbilicus.

Discussion. Parviscala lacinia sp. nov. is highly reminiscent of the extant Mediterranean Parviscala vittata ( Jeffreys, 1884) , which is only slightly larger (SL: 7 mm). It differs from the Paratethyan species in its slenderer shell and more convex whorls (see Weil et al. 1999: fig. 108; Bouchet & Warén 1986: figs 1193–1194). The protoconch of Parviscala vittata agrees with that of P. lacinia in its high conical outline with slightly angled last protoconch whorl (see Bouchet & Warén 1986: fig. 1174; https://www.gruppomalacologicoscalaria.org/atlas/epitonium-vittatumjeffreys-1884/). The extant Atlanto-Mediterranean Parviscala algeriana ( Weinkauff, 1866) , the type species of Parviscala , has a much broader shell, more lamellae and weaker spines (see Bouchet & Warén 1986: figs 1188– 1190). The extant eastern Pacific Parviscala deifica ( Melvill & Standen, 1903) and Parviscala eusculpta (Sowerby, 1903) are both larger and differ in the greater number of lamellae ( P. eusculpta ) or the wider apical angle ( P. deifica ) (see Weil et al. 1999: figs 316, 317; Brown & Raines 2004: fig. 1). Parviscala thyraea ( Kilburn, 1985) , from South Africa, differs from P. lacinia in its weak shoulder angulation and much weaker spines (see Kilburn 1985: fig. 139; Weil et al. 1999: fig. 169). Parviscala climacotum ( Kilburn, 1985) and P. columba ( Kilburn, 1985) , both extant species from South Africa, differ from the Paratethyan species in their broader shells and fewer lamellae with weaker spines (see Kilburn 1985: figs 133, 136; Weil et al. fig. 163).

Paleoenvironment. The common occurrence at Steinebrunn ( Austria) suggests shallow marine environments with sea grass (own data M.H.).

Distribution in Central Paratethys. Badenian (Middle Miocene): Polish-Ukrainian Fore-Carpathian Basin: Shushkivtsi (= Szuszkowce) ( Ukraine) ( Friedberg 1928); Vienna Basin: Steinebrunn ( Austria) (hoc opus); Pannonian Basin: Hidas, Szokolya, Várpalota,( Hungary) ( Strausz 1966); Southern Pannonian Basin: Jazovac, Hrvaćani ( Bosnia and Herzegovina) ( Atanacković 1985); Făget Basin: CoŞteiu de Sus ( Romania) (hoc opus).