Tapiena munae, Tan & Damit & Japir & Chung & Robillard, 2025

Tan, Ming Kai, Damit, Dayang Fazrinah Binti Awg, Japir, Razy, Chung, Arthur Y. C. & Robillard, Tony, 2025, New species, new locality records and descriptions of calling songs of Phaneropterinae from Sabah, Zootaxa 5604 (4), pp. 505-528 : 523-527

publication ID	https://doi.org/10.11646/zootaxa.5604.4.4
publication LSID	lsid:zoobank.org:pub:43CDF92A-7B39-4926-8FC0-EA813B85DF0E
DOI	https://doi.org/10.5281/zenodo.15214101
persistent identifier	https://treatment.plazi.org/id/03D78787-A50D-4C1D-8DC5-FDEDFDA9FF12
treatment provided by	Plazi
scientific name	Tapiena munae
status	sp. nov.

Treatment

Tapiena munae View in CoL sp. nov.

( Figs 19–21 View FIGURE 19 View FIGURE 20 View FIGURE 21 )

Material examined. Holotype. EAST MALAYSIA; Sabah State • 1♂; Sinipung Hill, near Long Pasia and Meligan ; N4.44538 E115.71494, 1556.0±10.1 m.a.s.l.; 21 November 2024, 21h30; attracted to light trap; coll. M.K. Tan; SBH.24.204 ( FRC). GoogleMaps

Paratype. EAST MALAYSIA; Sabah State • 1♂; Sinipung Hill, near Long Pasia and Meligan ; N4.43477 E115.72321, 1290.0±7.0 m.a.s.l.; 20 November 2024, 19h36; on foliage of small tree; coll. M.K. Tan; SBH.24.181 ( ZRC) GoogleMaps .

Diagnosis. The new species has a stout habitus and is characterised by the following characters: its face is red, from the fastigium to the frons, clypeus and labrum; the male tenth abdominal tergite forms a simple plate with two triangular lateral lobes (with rounded apices) and has its posterior margin between the lateral lobes truncated and roundly excised in the middle; the cercus is stout, bent at the middle (instead of arcuated) with its apex having three tooth-like spines.

The new species is generally similar to T. paraincisa , Tapiena?incisa and Tapiena dimidia sp. nov. owing to its stout habitus and its fore femur and tympanum red and the simple tenth abdominal tergite; but it differs in all the characters described in the previous paragraph. The new species also differs from T. paraincisa by the stridulatory file more densely packed with teeth (at the middle with circa 82 teeth per mm instead of 52 teeth per mm).

Etymology. This species is named after Munchie ‘Mun’, the first author’s dog who passed away on 15 November 2024.

Description. Habitus as shown in Figs 19 View FIGURE 19 , 20 View FIGURE 20 ; generally typical for genus and similar to species Tapiena?incisa and Tapiena dimidia sp. nov. Fastigium in dorsal view narrowly triangular, short, feebly surpassing anterior apex of eyes; furrowed longitudinally, lateral of furrow swollen; sulcus as wide as antennal scapus; with apex roundly truncated ( Fig. 21A View FIGURE 21 ). Eye prominent, in dorsal view globular slightly protruding anteriorly ( Fig. 21A View FIGURE 21 ); median and lateral ocelli prominent, round and large ( Fig. 21C View FIGURE 21 ). Frons very densely and strongly punctuated ( Fig. 21C View FIGURE 21 ); dorsum of head ( Fig. 21A View FIGURE 21 ) and genae ( Fig. 21B View FIGURE 21 ) densely punctuated. Pronotum circa 1.1 times longer than wide, densely punctuated ( Figs. 21A View FIGURE 21 ). Dorsal disc with anterior margin straight; lateral margins straight and nearly parallel (feebly diverging posteriorly); posterior margin broadly rounded; hind lobe after middle, separated by indistinct and shallow suture; longitudinal furrow barely visible ( Fig. 21A View FIGURE 21 ). Lateral keel present. Lateral lobe 1.2 times as tall as long; ventral margin fairly long and straight, anterior and posterior margins substraight ( Fig. 21B View FIGURE 21 ). Thoracic auditory spiracle [= thoracic foramen] covered by pronotal lateral lobe ( Fig. 21B View FIGURE 21 ). Prosternum unarmed. Mesosternum triangular with subacute apex; metasternum rectangular with anterior angle obtusely rounded, posterior angle 90° and rounded. Tegmen and hind wing fully developed. Hind wing with apex distinctively projecting beyond tegmen; exposed apex with dorsal and ventral margins straight and tapering to subacute apex ( Figs 20C, 20D View FIGURE 20 ). Legs generally with very short and fine setae. Fore coxa with distinct, long, fairly straight (only slightly curved at base) spine. Fore tibia with external tympanum open, oval; internal tympanum slit covered by large conchate swelling ( Fig. 21D View FIGURE 21 ). Hind femur with genicular lobe rounded, without any spine.

Male. Tegmen with anterior margin straight and posterior margins convex, distinctly widened in the middle before tapering to narrowly-rounded apex; punctuated with many small transparent cells ( Figs 20A, 20B View FIGURE 20 ). Tegminal venation: Radius sector (Rs) originating before mid-length, with fork longer than stem ( Figs 20A, 20B View FIGURE 20 ). Stridulatory file of left tegmen crescent shaped, 2.9 mm in length ( Fig. 21E View FIGURE 21 ); area around file not swollen; file at anal end with circa 16 stout teeth, thereafter, teeth very densely and evenly spaced elongated (more closely packed at the sides than in the middle); file at middle with circa 82 teeth per mm and teeth circa 0.11 mm long ( Fig. 21E View FIGURE 21 ). Mirror on left tegmen made up of network of inter-crossed veins, margins unclear ( Fig. 21E View FIGURE 21 ). Mirror on right tegmen longer than broad, rectangular but somewhat angularly bent at middle ( Fig. 21F View FIGURE 21 ).

Tenth abdominal tergite simple, with little or no modification, without tooth or long process apically; at posterior end with two triangular lateral lobes (with rounded apices) and having posterior margin between lateral lobes truncated and roundly excised at middle ( Figs 21G–I View FIGURE 21 ). Epiproct tongue-shaped, concealed beneath tenth abdominal tergite ( Fig. 21I View FIGURE 21 ). Cercus stout, bent at middle (instead of arcuated) with its apex having three tooth-like spines; two lateral spines larger than medial one ( Figs 21G–I View FIGURE 21 ). Subgenital plate broad basally, gently tapering into shaft; with anterior margin concave; with posterior margin deeply excised at middle, laterally extended into two long and narrow lobes; styli slightly longer than apical excision of subgenital plate, tapering into obtuse apex ( Fig. 21J View FIGURE 21 ).

Female. Unknown.

Colouration. Generally green when alive ( Fig. 19 View FIGURE 19 ), except face red, from fastigium to frons to clypeus and labrum (except median ocellus); scape and pedicel black dorsally, red ventrally; apical segment of maxillary palps and lateral keels of pronotum having tints of red ( Figs 21A–C View FIGURE 21 ). Fore coxa (apical end), fore femur and anterior half of fore tibia (including tympanum) red with tint of black ( Fig. 21D View FIGURE 21 ). Abdomen mostly white or pale green when alive ( Fig. 19 View FIGURE 19 ).

Measurements (in mm). See Table 1 View TABLE 1 .