Sanganus quadrangularis, Kallenborn & Schmolke, 2017

Kallenborn, H. G. & Schmolke, F., 2017, Review of the genus Sanganus Distant, 1909 including the description of a new species, S. quadrangularis spec. nov. (Hemiptera, Heteroptera, Tessaratomidae), Spixiana 40, No. 1, pp. 75-88 : 86-87

Sanganus quadrangularis spec. nov.

Fig. 1C

Type material. Holotype ♀ ( ZSM): “Philippinen [handwritten], leg. G. Böttcher [printed] / Pseudopycanum quadrangularis [handwritten]” [ Philippines, Luzon, Samar or Mindanao?; 1914-1919]. Remark: Georg Böttcher (1890-1920), a German insect-trader, spent several years between 1914 and 1919 in the Philippines ( Weidner 1964), so that the specimen must be dated from this period. Unfortunately, the precise locality is not specified, but Böttcher is known to have collected in Luzon, Prov. Laguna (possibly his dwelling place), Samar, and Mindanao. The species label was probably written by Hans Werner Taeuber, an antiquarian and heteropterologist from Munich, who came into possession of the vast majority of the Heteroptera collected by Böttcher (now kept in the ZSM). Neither a valid name, nor a description of the specimen has ever been published. – Paratype ♀ (cJPM): “Philippines, Gutalac, Zamboanga del Norte, Mindanao, XI 2014, I. Lumawig [handwritten]” .

Description

Colouration. Entire body of dried specimens reddish to saddle brown.

Head. Mandibular plates subtriangular, distally acute ( Fig. 2 C). Antennal segments obscure ( Fig. 3 C).

Pronotum. Lateral margins nearly straight and parallel, margins fuscous laterally and antero-laterally ( Fig. 4 C); punctures fine and concolourous ( Fig. 4.1 C); humeral angles distally not surpassing head.

Scutellum. Surface wrinkled; punctures concolourous, fine and densely arranged ( Fig. 5 C); distal tongue concolourous, scarcely punctate.

Hemelytra. Corium with fading venation, lateral margin entirely slightly convex; membrane distally broadly rounded, not extending over abdomen ( Fig. 6 C).

Sternum. Metasternal carina sulcate along the entire median length of the sternum ( Fig. 7 C).

Metathoracic scent efferent system. Mesopleural evaporatorium reduced to a narrow strip, not widened apically, basally connected with posterior mesopleural margin ( Fig. 8 C).

Abdomen. As in generic description ( Fig. 10 C).

Male genitalia. Not available for this study. Kumar & Ghauri (1970, figure 68) illustrated the paramere of a species named “ Pseudopycanum quadrangularis Taeuber ”, without providing any details in the text. The fate of this male individual, which may have been in their keeping, could not be clarified. We reproduce this figure with all due caution (see Fig. 12 C). It shows a less strongly bent apophysis of the paramere and a moderately protruding sensory lobe.

Female genitalia. Proximo-distal angle of laterotergites VIII spinose. Laterotergites IX subquadrangular, lateral margins concave, sutural margins distally diverging and straight ( Fig. 14 C). Spermatheca as in generic description; combined length of bulb and pumping region (1.4 mm) slightly smaller than in S. westwoodii (1.7 mm) and S. jenseni (1.6 mm); cup-like region of distal flange more extended ( Fig. 15 C).

Measurements of the ♀ holotype and ♀ paratype (in parentheses). Total body length: 31.7 mm (34.5 mm). Maximum body width across abdomen: 19.6 mm (21.5 mm). Head width: 4.4 mm (4.8 mm). Ratios of antennal segments I to IV: 0.22:0.68:0.61:1.00 (0.21:0.76:0.69:1.00).Maximum pronotum width: 17.7 mm (18.6 mm). Maximum pronotum length: 8.4 mm (9.1 mm). Median scutellum length: 7.1 mm (8.1 mm). Basal scutellum width: 10.7 mm (12.2 mm).

Etymology. The species name quadrangularis (= is, = e) is a Latin adjective composed of the words quattuor = four and angulus = edge, angle, and refers to the nearly rectangular body shape.

Distribution ( Fig.17). Philippines: Mindanao (Zamboanga del Norte).

Comment. Although the pronotal shapes appear similar, we do not share Taeuber’ s opinion (original label of the holotype) that this species belongs to Pseudopycanum ( Fig. 1 D) for the reasons mentioned in the generic key.