Polyosma affinis Schulze-Menz ex B.J.Conn & O.K.Paul, 2024

Paul, O. K., Conn, B. J. & Henwood, M. J., 2024, Taxonomic review of Polyosma (Escalloniaceae) in Papuasia, Blumea 69, pp. 54-88 : 63-65

KEY TO THE SPECIES OF POLYOSMA IN PAPUASIA

1. Leaf margin entire.Although leaf margin usually entire, often with 1 or 2 poorly developed teeth distally............ 2

1. Leaf margin toothed........................... 17

2. Inflorescence pendulous at flowering........ 5. P. buxea

2. Inflorescence erect at flowering.................... 3

3. Leaves (when mature) with abaxial surface hairy...... 4

3. Leaves with abaxial surface glabrous, or with an occasional hair.......................................... 7

4. Leaf lamina up to 5 by 1–2 cm, with 4–7 lateral veins on each side of midvein; petiole less than 1 cm long. — Although leaf margin usually entire, often with 1 or 2 poorly developed teeth distally.................. 20. P. longebracteolata

4. Leaf lamina greater than 7 by 2–8 cm, with at least 10 lateral veins on each side of midvein; petiole usually at least 1 cm long......................................... 5

5. Leaf base rounded; leaves 2–3.5 cm wide; corolla buds brown with green tinge........... 28. P. schulzemenzii

5. Leaf base acute; leaves 3.5–8 cm wide; corolla buds white to pale yellow or pinkish green to purplish............ 6

6. Leaf lamina elliptic, 9–11 by 3.5–4.5 cm; secondary veins of lamina 10–15 on each side of midrib; petiole 0.8–1.5 cm long............................ 26. P. pubescens

6. Leaf lamina elliptic to narrowly obovate, 11–25 by (4–) 5– 8 cm; secondary veins of lamina 17–28 on each side of midrib; petiole 1.5–2(–2.8) cm long.. 18. P. kamialiensis

7. Leaf lamina ± obovate........................... 8

7. Leaf lamina elliptic or narrowly ovate............... 10 8. Leaf lamina up to 2.7 by 0.5–1.1 cm. — Leaf margin rarely entire, usually shortly serrate, with 1 or 2 teeth per side................................... 27. P. rampae

8. Leaf lamina at least 4 by 1.3 cm .................. 9

9. Leaf lamina 5–12 by 3–7 cm; petiole 1.5–3 cm long................................... 12. P. glaberrima

9. Leaf lamina 4–6 by 1.3–2 cm; petiole 0.7–1 cm long................................. 30. P. stenosiphon

10. Venation usually prominent on abaxial surface of lamina........................................... 11

10. Venation usually indistinct on abaxial surface of lamina. ... ... ... ... ... ... .. ... ... ... ... ... ... .... 13

11. Leaf apex attenuate; petiole 5–25 cm long............................................ 6. P. cestroides

11. Leaf apex obtuse to rounded or acuminate; petiole 0.5– 3 cm long................................... 12

12. Leaf apex acuminate; lamina with 13–42 lateral veins on each side of midvein; inflorescence (9–) 10–22 cm long; corolla 5–10 mm long.............. 17. P. integrifolia

12. Leaf apex obtuse or rounded; lamina with c. 10 lateral veins on each side of midvein; inflorescence up to 5 cm long; corolla up to 28 mm long............. 24. P. oligantha

13. Leaf lamina with apex long-tapering or attenuate.................................... 3. P. amygdaloides

13. Leaf lamina with apex subacute or acuminate...... 14

14. Inflorescence glabrous........... 21. P. macrobotrys

14. Inflorescence sparsely hairy.................... 15

15. Branchlets sparsely hairy; subcanopy tree, up to c. 5 m high............................... 9. P. flenleyi

15. Branchlets glabrous; canopy or subcanopy tree, c. 10– 20 m high................................... 16

16. Branchlets with white lenticels; corolla 8–12 mm, green, brownish green, sometimes becoming brownish maroon; leaf blades 4– 9 cm long.................1. P. affinis

16. Branchlets with prominent craterous scars; corolla 3–5 mm long, whitish yellow; leaf blades 5.5–11 cm long.........................................11. P. gigantea

17. Inflorescence axillary or subterminal (appearing axillary). ... ... ... ... ... ... .. ... ... ... ... ... ... .... 18

17. Inflorescence distinctly terminal................. 25

18. Bracteoles and calyx lobes glabrous.... 8. P. finisterrae

18. Bracteoles and calyx lobes hairy................. 19

19. Leaf lamina with apex mucronate....... 23. P. occulta

19. Leaf lamina with apex acuminate or attenuate...... 20

20. Venation of abaxial leaf lamina visible but usually not prominent...................... 16. P. infernaralis

20. Venation of abaxial leaf lamina usually prominent... 21

21. Leaf lamina glabrous.................. 7. P. dentata

21. Leaf lamina hairy............................. 22

22. Leaf lamina rhombic-oblong or narrowly obovate..................................... 13. P. heliciiformis

22. Leaf lamina elliptic or broadly elliptic.............. 23

23. Calyx lobes > 5 mm long; bracteoles> 5 mm long....................................... 10. P. forbesii

23. Calyx lobes <5 mm long; bracteoles <5 mm long... 24

24. Leaf lamina densely ferruginous; 6–14 by 3–6 cm ......................................... 4. P. brassii

24. Leaf lamina sparsely hairy; 5–22 by 2–7 cm ......................................... 14. P. helicioides

25. Leaf lamina hairy, at least on abaxial surface....... 26

25. Leaf lamina glabrous.......................... 30

26. Leaf apex attenuate; lamina broadly elliptic........ 27

26. Leaf apex usually obtuse to acuminate; lamina oblong, elliptic, obovate to spathulate................... 28

27. Subcanopy tree, 10–20 m tall; bracteoles 5–10 mm long.................................. 10. P. forbesii

27. Small tree, c. 3 m tall; bracteoles 1–2 mm long.......................................... 15. P. induta

28. Leaf lamina 0.2–0.5 by 0.5–1.8 cm, oblong, or obovate to spathulate; apex obtuse to subacute; bracteoles (3–4 mm long) as long as calyx. — Leaf margin is usually entire, but often with 1 or 2 poorly developed teeth........................................ 20. P. longebracteolata

28. Leaf lamina 13.5–17 by 4–8 cm, obovate or elliptic; apex obtuse or acuminate; bracteoles either shorter than (2–3 mm long) or mostly longer (5–10 mm long) than calyx...................................... 29

29. Leaf apex subacute to acute; bracteoles 5–10 mm long (mostly longer than calyx)............. 2. P. alipensis

29. Leaf apex acuminate; bracteoles 2–3 mm long (shorter than calyx).................... 25. P. pachyrhachis

30. Venation of abaxial surface of lamina usually prominent ........................................... 31

30. Venation of abaxial surface of lamina visible but usually not prominent............................... 36

31. Shrub c. 2 m tall; inflorescence with rachis 1.5–2 cm long; leaf lamina up to 2.7 cm long.......... 27. P. rampae

31. Small tree up to 5 m tall, or large tree, 6–20 m tall; inflorescence with rachis at least 4 cm long; leaf lamina at least 3.5 cm long................................. 32

32. Leaf lamina obovate-narrowly ovate or elliptic-narrowly ovate; leaf apex obtuse to apiculate (on distal leaves)................................. 33. P. trimeniifolia

32. Leaf lamina elliptic; leaf apex acuminate.......... 33

33. Leaf margin distinctly serrate throughout; corolla purple with a reddish or greenish tinge.... 29. P. scyphocalyx

33. Leaf margin with an occasional tooth; corolla buds white or yellow................................... 34

34. Corolla buds white................. 17. P. integrifolia

34. Corolla buds whitish yellow, often with green tinge... 35

35. Leaf lamina with 13–25 lateral veins on each side of midvein; corolla buds 3–5 mm long; rhachis 8.5–10 cm long................................. 11. P. gigantea

35. Leaf lamina with c. 8 lateral veins on each side of midvein; corolla buds c. 10 mm long; rhachis 5–6 cm long.................................... 32. P. torricellensis

36. Leaf apex acuminate or attenuate....19. P. leptorhachis

36. Leaf apex mucronate.......................... 37

37. Shrub up to 1 m tall; corolla green. — Montane forest (at elevation of up to 3500 m)......... 22. P. mucronata

37. Small tree, 2–5 m tall; corolla greenish brown or purple. — Alpine shrubbery forest (from elevations of at least 3000 m)......................... 31. P. subalpina