Geosaurina Bonaparte, 1845

Young, Mark T., Wilberg, Eric W., Johnson, Michela M., Herrera, Yanina, Brandalise, Marco de Andrade, Brignon, Arnaud, Sachs, Sven, Abel, Pascal, Foffa, Davide, Fernández, Marta S., Vignaud, Patrick, Cowgill, Thomas & Brusatte, Stephen L., 2024, The history, systematics, and nomenclature of Thalattosuchia (Archosauria: Crocodylomorpha), Zoological Journal of the Linnean Society 200 (2), pp. 547-617 : 607

Geosaurina Bonaparte, 1845 (Zoological Code)

Geosaurina Foffa et al. 2018: 1125, converted clade name (PhyloCode)

RegNum registration number 1022.

Type genus: Geosaurus Cuvier, 1824 .

Etymology

‘Pertaining to Geosaurus ’. The stem Geosaur - is based on the genus Geosaurus . - ina, is a Latin neuter plural suffix for ‘pertaining to’. Under Article 29.2 of the Zoological Code, the suffix - ina denotes a subtribe rank within the family-group.

Geological range

Middle Jurassic (middle Callovian) to Early Cretaceous (late Valanginian) ( Debelmas and Strannoloubsky 1956, Foffa et al. 2018).

PhyloCode phylogenetic definition

The largest clade within Metriorhynchidae containing Geosaurus giganteus ( von Sömmerring 1816) , but not Metriorhynchus brevirostris ( Holl 1829) , Plesiosuchus manselii ( Hulke 1870) , Torvoneustes carpenteri ( Wilkinson et al. 2008) , and Dakosaurus maximus ( Plieninger 1846) .

Reference phylogeny

Fig. 3 .

Zoological Code diagnosis and PhyloCode diagnostic apomorphies

Metriorhynchid crocodylomorphs with the following unique combination of characters (5): maxilla ornamented with a conspicuous-to-faint pitted pattern (18.1); presence of apicobasal facets on the labial surface of the tooth crowns (within Thalattosuchia shared with Cricosaurus puelchorum ) (639.1); presence of laminar tooth crowns (within Thalattosuchia shared with Cricosaurus puelchorum ) (640.1); maxillary overbite of the dentary dentition at the mid-dentition (660.1) and posterior dentition (661.1).

Composition

Ieldraan and Geosaurus .

Comments

Authorship: The nomen Geosaurina was first used by Foffa et al. (2018), however under the Zoological Code Bonaparte 1845 is the nominal authority. The nominal author of a family-group is the author who first erected a family-group taxon that is valid (in fulfilment of Article 11), and in accordance with the Zoological Code Principle of Coordination applied to family-group names (Article 36.1). Under the PhyloCode, Foffa et al. (2018) is the nominal authority.

Prior phylogenetic definition: Foffa et al. (2018) did not phylogenetically define Geosaurina .

Figures

Figure 3. The equal weights, no constraints strict consensus topology (C0). Length = 4164, CI = 0.303, RI = 0.831, RC = 0.251, HI = 0.697, number of most parsimonious cladograms hit the hold limit.Phylogeny pruned to highlight Thalattosuchia (see Supporting Information, Appendix S2, Fig. S1 for the entire topology). A, Crocodylomorpha; B, Crocodyliformes; C, Thalattosuchia; D, Neothalattosuchia; E, Teleosauridae; F, Teleosaurinae; G, Aeolodontini; H, Machimosauridae; I, Machimosaurinae; J, Machimosaurini; K, Euthalattosuchia; L, Metriorhynchidae; M, Metriorhynchinae; N, Rhacheosaurini; O, Geosaurinae; P, Geosaurini. Green lines highlight non-neothalattosuchian thalattosuchians, orange lines highlight teleosauroid thalattosuchians, and blue lines highlight metriorhynchoid thalattosuchians.Images from PhyloPics.org, with the teleosauroid by Scott Hartman, and the metriorhynchid by Dmitry Bogdanov (vectorized by T. Michael Keesey), the only change made was to their colouration, and their license is: https://creativecommons.org/licenses/by/3.0/.