Beremendia fissidens ( Petényi, 1864 )

Botka, D. & Mészáros, L., 2014, (Mammalia, Soricidae) remains from the late Early Pleistocene Somssich Hill 2 locality (Southern Hungary) and their taxonomic, biostratigraphical, palaeoecological and palaeobiogeographical relations, Fragmenta Palaeontologica Hungarica 31, pp. 83-115 : 86-87

Beremendia fissidens ( Petényi, 1864)

( Figs 2–12 , 16 )

1864 Crossopus fissidens n. gen. n. sp. – PETÉNYI, pp. 60–70, pl. I, fig. 5.

1911 Neomys fissidens (Petényi) – KORMOS, pp. 156–158, pl. VII, figs 1–3.

1930 Neomys (?) fissidens (Petényi) – HELLER, pp. 254–258, text-figs 2–4, pl. XV, figs 1–3.

1930 Beremendia fissidens n. gen. (Petényi) – KORMOS, pp. 43–44, 57.

1934 Beremendia fissidens (Petényi) – KORMOS, pp. 299–301, fig. 33.

1936 Beremendia fissidens (Petényi) – HELLER, pp. 107–108, pl. VII, figs 1–2.

1949 Beremendia fissidens (Petényi) – FRIANT, pp. 256–257, fig. 17.

1958 Beremendia fissidens (Petényi) – KOWALSKI, pp. 13–14, fig. 4.

1959 Beremendia fissidens (Petényi) – SULIMSKI, pp. 152–154, tab. 7, text-fig. 4: 1a-f, pl. III, fig. 7.

1966 Beremendia fissidens (Petényi) – FEJFAR, figs 4f-j, 5f-h, 10d, h.

1967 Beremendia fissidens (Petényi) – REPENNING, pp. 50–51, fig. 35.

1976 Beremendia fissidens (Petényi) – FREUDENTHAL et al., pl. I. figs 7–8.

1976 Beremendia fissidens (Petényi) – RZEBIK-KOWALSKA, pp. 360–368, tabs I–III, X, figs 1–11, 21, 23, 25, 28–29, 33–35.

1979 Nectogalinia altaica n. gen. n. sp. – GUREEV, p. 458, fig. 235.

1984 Beremendia fissidens (Petényi) – REUMER, pp. 131–138, text-fig. 16, pl. 32, figs 1–13, pl. 33, figs 1–6, pl. 34, figs 1–4, pl. 35, figs 1–2.

1996 Beremendia fissidens (Petényi) – DAHLMANN & STORCH, p. 185, tab. 3.

2002 Beremendia fissidens (Petényi) – AGUILAR et al., p. 24, pl. II, figs 11–14.

2003 Beremendia fissidens (Petényi) – POPOV, pp. 60–62, tab. V, figs 7–8.

2004 Beremendia fissidens (Petényi) – SUÁREZ & MEIN, pp. 122–123, fig. 8c-e.

2006 Beremendia fissidens (Petényi) – HARRISON et al., pp. 121–124, figs 1a-c, 3c, 4a-b.

2007 Beremendia cf. fissidens (Petényi) – ČERMÁK et al., fig. 3a, c.

2007 Beremendia fissidens (Petényi) – CUENCA-BESCÓS & ROFES, pp. 113–115, fig. 3a.

2007 Beremendia fissidens (Petényi) – HORÁČEK et al., pp. 459–460, pl. 2, figs 3–4.

2009 Beremendia fissidens (Petényi) – ROFES & CUENCA-BESCÓS, pp. 24–30, tab. 1, figs 3–5. 2010 Beremendia fissidens (Petényi) – FURIÓ et al., pp. 929–936, figs 2–8.

2012 Beremendia fissidens (Petényi) – HELLMUND & ZIEGLER, p. 81, pl. 1, fig. 3.

2013 Beremendia fissidens (Petényi) – CUENCA-BESCÓS et al., fig. 4d.

2013 Beremendia fissidens (Petényi) – RZEBIK-KOWALSKA, pp. 25–28, tabs 30–32, fig. 5: 7. 2014 Beremendia fissidens (Petényi) – BENNÀSAR et al., tab. 2, figs 3–4.

in press Beremendia fissidens (Petényi) – BOTKA & MÉSZÁROS, tabs 1–2, pl. I, figs 1–7.

Type material – Crossopus fissidens Petényi, 1864 : Hungarian Natural History Museum , Department of Palaeontology and Geology, inventory number V 61.1585 . Syntypes: 1 skull fragment, 2 left and 2 right maxillae, 13 left and 15 right mandibles, 1 condyle and 8 I

1

.

Type locality – Beremend limestone quarry (locality2), Villány Hills, Hungary, Upper Pliocene (Villányian Stage, Beremend Phase, MN 16 ) .

Studied material – Somssich Hill 2 locality: 13 left maxillary fragments, 11 right maxillary fragments, 9 left mandibles, 6 right mandibles, 20 left mandible fragments, 18 right mandible fragments, 8 left I 1, 13 right I 1, 5 left P 4, 6 right P 4, 5 left M 1, 4 right M 1, 3 left M 2, 2 right M 2, 10 left I 1, 5 right I 1, 1 left A 1, 1 left A 2, 8 left M 1, 6 right M 1, 3 left M 2, 7 right M 2, 1 right M 3 and 4 humeri.

Measurements – Table 1.

Dental formula – 153/123 or 143/123 (A 4 is missing from a few specimens).

Figures

Figs 2–3. Beremendia fissidens (Petényi, 1864), Somssich Hill 2. 2 = left maxillary fragment with P4-M2, layer 14, occlusal view, scale bar: 1 mm, 3 = left mandible with I -M , layer 30, buccal view, 1 2 scale bar: 2 mm

Figs 4–6. Beremendia fissidens (Petényi,1864), Somssich Hill 2, scale bar:2 mm.4 = right mandible fragment with I 1 , A 2 and M 1 , layer 13, lingual view, 5 = right mandible fragment with M 2 -M 3 , layer 28, lingual view, 6 = left mandible fragment with M -M , layer 32, occlusal view

Figs 7–9. Beremendia fissidens (Petényi, 1864), Somssich Hill 2, scale bar: 1 mm. 7 = left M 3, layer 32, occlusal view, 8 = left M , layer 5, occlusal view, 9 = left I1, layer 11, buccal view

Figs 10–12. Beremendia fissidens (Petényi, 1864), Somssich Hill 2, scale bar: 1 mm. 10 = left maxillary fragment with P4-M1, layer 32, occlusal view, 11 = right M1, layer 28, occlusal view, 12 = left maxillary fragment with M2, layer 31, occlusal view

Figs 15–16. Beremendia fissidens (Petényi, 1864) and Beremendia minor Rzebik-Kowalska, 1976, Somssich Hill 2, scale bar: 0.5 mm. 15 = B. minor right M3, layer 35, occlusal view, 16 = B. fissidens right I 1, layer 13, medial view, the venom groove is marked by the white arrow