Arisemus Satchell, 1955

Durán-Luz, Juana & Ibáñez-Bernal, Sergio, 2025, Two new species of Arisemus Satchell, and first records of A. atrasetus (Rapp) and A. salazari Quate from Mexico (Diptera, Psychodidae), Zootaxa 5618 (3), pp. 411-423 : 412

Arisemus Satchell

Telmatoscopus (Arisemus) Satchell, 1955: 88 .

Arisemus Satchell : Botosaneanu & Vaillant, 1970: 176; Duckhouse, 1974: 149; Vaillant, 1986: 334; Quate, 1996: 21; Quate & Brown, 2004: 71.

Syn. Bazara Vaillant, 1986: 337 ( Quate & Brown 2004) .

In general, the diagnostic characters provided by Quate & Brown (2004) and followed by Wagner & Ibáñez-Bernal (2009) in their key for identification, allows Arisemus to be distinguished from other psychodine genera. Nevertheless, it is important to add that some species of Arisemus present scent organs on the anepisternum, whose shape has been shown to vary according to the species, and that the scape may be slightly longer than the pedicel or exceeding up to three times its length.

In recent years, changes have been made to the nomenclature of male terminalia structures with the intention to recognizing homologies, while the observation of them have been done in more detail. Some suggestions were made by Curler & Moulton (2012), Kvifte (2014) and Kvifte & Wagner (2017). Male terminalia of Arisemus species are characterized as follows: anterior gonocoxal condyles (formerly anterior gonocoxal apodemes, inner condyles) expanded, connected to the ejaculatory apodeme (formerly basiphallus, aedeagal apodeme) by a median keel; ejaculatory apodeme large, spoon- or paddle-shaped; gonostylus simple or with two or three branches; epandrium (tergite IX) with one or two foramina; hypopod (formerly surstylus, cercopod, epandrial appendage) with one tenaculum (sometimes referred as retinaculum) at apex; hypandrium short (interpreted as “gonocoxites broadly fused at base”) or long and belt-like (interpreted as “gonocoxites connected by hypandrium”); epiproct (X tergite) and hypoproct (X sternite) tongue-like. Regarding the aedeagus, Quate & Brown (2004) mentioned it as asymmetrical; nevertheless, there are species in which it can be nearly symmetrical, as in Arisemus imeldae Ibáñez-Bernal & Suárez-Landa, 2017 , to large and asymmetrical, as in Arisemus amydrus Quate & Brown, 2004 ; parameres sometimes longer than aedeagus, but with a membranous parameral sheath dorsal to aedeagus. It is highly probable that the parameral sheath has not been documented in several species due to the maceration procedure of specimens, but it may be present at base of the aedeagus, a character that is shared with Maruina Müller , where it is well represented, and that could strengthen the grouping of the tribe Maruinini Enderlein , as suggested by the most recent proposal of tribal classification ( Kvifte 2018).